Phylogenetic analysis of the Neotropical scarab beetle tribe Aegidiini (Coleoptera, Scarabaeidae, Orphninae) with description of new taxa Author Frolov, Andrey V. https://orcid.org/0000-0002-6724-6828 Laboratory of Insect Systematics, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab., 1, St. - Petersburg 199034, Russia afrolov@zin.ru Author Akhmetova, Lilia A. https://orcid.org/0000-0002-2151-1278 Laboratory of Insect Systematics, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab., 1, St. - Petersburg 199034, Russia Author Neita-Moreno, Jhon Cesar https://orcid.org/0000-0003-2998-2063 Instituto de Investigaciones de Recursos Biologicos Alexander von Humboldt, Claustro de San Agustin, Boyaca, Colombia text ZooKeys 2023 2023-06-06 1166 33 47 http://dx.doi.org/10.3897/zookeys.1166.102813 journal article http://dx.doi.org/10.3897/zookeys.1166.102813 1313-2970-1166-33 94B60F4A84304FDA9C921B9C66833E83 1E6AF92F558B59BFBC9CAB594DB056A6 Subtribe Aegidiina Paulian, 1984 Type genus. Aegidium . Diagnosis. Small to medium-sized beetles (body length 5-20 mm), brown to black colored without pattern, more or less densely punctate, smooth or densely setose. Mandibles subsymmetrical, without lateral processes, distinctly or feebly protruding past anterior margin of frontoclypeus in dorsal view. Labrum exposed or hidden under clypeus in dorsal view. Frontoclypeus symmetrical or subsymmetrical, without tubercles, horns or ridges, or in males with variably shaped bilobate anterior frontoclypeal process. Pronotum of males may be with deep excavation in the middle, with 2 horns or ridges bordering the excavation near anterior margin (lateral pronotal processes), and with a tubercle or small horn medially on the anterior margin (anterior pronotal process); these characters are subject to allometric variability and may not be developed in some males. Females have a convex pronotum without armature or pronotum impressed anteriorly on disc and with a tubercle medially on anterior margin. Propleurae with carinae separating anterolateral areas from basal area. Scutellum narrowly rounded apically, about 1/8-1/13 length of elytra. Elytra convex, with marked humeral umbones (except for brachypterous species). Surface flat or with two low ridges in basal half; the ridges may be more or less convex, smooth, to almost indistinct. Pubescence of dorsal side indistinct or dense. Wings fully developed or vestigial. Metepisternon triangular, its posterior angle rounded to triangular and situated in distinct concavity of epipleuron. Mesocoxal cavities connected by a hole. Protibiae with three outer teeth, somewhat serrate basad of the teeth, with a smaller, medial tooth in majority of males. In males, anterior spur is absent. Each procoxae with one elongate hollow. Mesotibiae with or without a tuft of setae ventroapically in males. Stridulatory file with relatively fine, evenly spaced carinae. Phallobase tube shaped with strongly sclerotized ventral side but without differentiation of ventral and dorsal sclerites; ventroapical plate absent or present. Parameres symmetrical, relatively long, apices tapering or curved downwards, with or without setae; a few species have complex, feeble sclerotised processes on the parameres lateroapically. Endophallus without armature or with a small group of spinules; in one species of Aegidium there is a sclerite with two large curved spines. Spiculum gastrale T-, Y- or V-shaped, with setae on apical plate. Subcoxites oval, with dense, long setae mediabasally; coxites triangular, long, with dense short setae mediabasally and sparse long setae apically; stili distinct, elongated, or not separated from coxites. Taxon composition. The subtribe is comprised of Aegidium Westwood, 1845 (25 spp), Paraegidium Vulcano et al., 1966 (6 spp), Aegidiellus Paulian, 1984 (3 spp) and Onorius Frolov & Vaz-de-Mello, 2015 (2 spp). Distribution. Endemic to South and Central America.