<document id="04865F06A73C77C52646108C46E1F4F2" ID-CLB-Dataset="28398" ID-DOI="10.11646/zootaxa.4565.2.12" ID-GBIF-Dataset="44ded8dc-3187-42db-a283-a59fec2c467d" ID-ISSN="1175-5326" ID-Zenodo-Dep="2589796" ID-ZooBank="9F4AA96D-877E-45EA-82FE-0DAFE856AD94" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552276784052" checkinUser="plazi" docAuthor="Chen, Zhi-Teng &amp; Wang, Bo" docDate="2019" docId="7B3787C7FF9F0B241880FCF23C0EFBE3" docLanguage="en" docName="zootaxa.4565.2.12.pdf" docOrigin="Zootaxa 4565 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Largusoperla acus Chen 2018" docType="treatment" docVersion="8" lastPageNumber="282" masterDocId="870EFFBFFF9E0B251817FFA63D70FFC1" masterDocTitle="Review of the fossil genus Largusoperla (Plecoptera: Perlidae): Annotated checklist, taxonomic identification, and description of a new species" masterLastPageNumber="291" masterPageNumber="281" pageNumber="282" updateTime="1698701381354" updateUser="ExternalLinkService">
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<mods:title id="00498610263D06B9C3977CB424E58A96">Review of the fossil genus Largusoperla (Plecoptera: Perlidae): Annotated checklist, taxonomic identification, and description of a new species</mods:title>
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<mods:namePart id="DCF9DEBAFB3F9BF551600CBCC7B4DE8A">Chen, Zhi-Teng</mods:namePart>
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<mods:title id="875FD0826EAB2395F7D7C969F4FD3D36">Zootaxa</mods:title>
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<mods:date id="B155158D89256AFDBF1A20653D88244E">2019</mods:date>
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<treatment id="7B3787C7FF9F0B241880FCF23C0EFBE3" ID-DOI="http://doi.org/10.5281/zenodo.5944768" ID-GBIF-Taxon="154366865" ID-Zenodo-Dep="5944768" LSID="urn:lsid:plazi:treatment:7B3787C7FF9F0B241880FCF23C0EFBE3" httpUri="http://treatment.plazi.org/id/7B3787C7FF9F0B241880FCF23C0EFBE3" lastPageNumber="282" pageId="1" pageNumber="282">
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<paragraph id="F32136D1FF9F0B241880FCF23F4AFCAF" blockId="1.[151,1436,852,1058]" box="[151,570,852,878]" pageId="1" pageNumber="282">
<heading id="A86981BDFF9F0B241880FCF23F4AFCAF" bold="true" box="[151,570,852,878]" fontSize="10" level="2" pageId="1" pageNumber="282" reason="2">
<taxonomicName id="349E4D52FF9F0B241880FCF23F4AFCAF" ID-CoL="6NXN6" authority="Chen et al., 2018" authorityName="Chen" authorityYear="2018" box="[151,570,852,878]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="282" phylum="Arthropoda" rank="species" species="acus">
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<emphasis id="C1EAEAC3FF9F0B241880FCF23C1EFCAC" bold="true" box="[151,366,852,877]" italics="true" pageId="1" pageNumber="282">Largusoperla acus</emphasis>
Chen 
<emphasis id="C1EAEAC3FF9F0B2419ABFCF03C8AFCAC" bold="true" box="[444,506,852,877]" italics="true" pageId="1" pageNumber="282">et al.,</emphasis>
2018
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<taxonomicName id="349E4D52FF9F0B241880FCDF3F3DFC53" authority="Chen et al. 2018" authorityName="Chen" authorityYear="2018" box="[151,589,889,914]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="282" phylum="Arthropoda" rank="species" species="acus">
<emphasis id="C1EAEAC3FF9F0B241880FCDF3C1FFC50" box="[151,367,889,913]" italics="true" pageId="1" pageNumber="282">Largusoperla acus</emphasis>
Chen 
<emphasis id="C1EAEAC3FF9F0B2419D2FCDC3F74FC50" box="[453,516,889,913]" italics="true" pageId="1" pageNumber="282">et al.</emphasis>
2018
</taxonomicName>
. Zootaxa, 4378(4): 575. Specimen status: male 
<typeStatus id="2C258873FF9F0B241C87FCDF3985FC53" box="[1168,1269,889,914]" pageId="1" pageNumber="282" type="holotype">holotype</typeStatus>
, NIGP167158 (NIGP).
</paragraph>
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<subSubSection id="BB84655AFF9F0B241880FC673C0EFBE3" pageId="1" pageNumber="282" type="discussion">
<paragraph id="F32136D1FF9F0B241880FC673C0EFBE3" blockId="1.[151,1436,852,1058]" pageId="1" pageNumber="282">
<emphasis id="C1EAEAC3FF9F0B241880FC673C7BFC1B" bold="true" box="[151,267,961,986]" pageId="1" pageNumber="282">Remarks.</emphasis>
<taxonomicName id="349E4D52FF9F0B241902FC673C9AFC18" box="[277,490,961,985]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="282" phylum="Arthropoda" rank="species" species="acus">
<emphasis id="C1EAEAC3FF9F0B241902FC673C9AFC18" box="[277,490,961,985]" italics="true" pageId="1" pageNumber="282">Largusoperla acus</emphasis>
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is the 
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species of 
<taxonomicName id="349E4D52FF9F0B241AE2FC673EFCFC18" box="[757,908,961,985]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="282" phylum="Arthropoda" rank="genus">
<emphasis id="C1EAEAC3FF9F0B241AE2FC673EFCFC18" box="[757,908,961,985]" italics="true" pageId="1" pageNumber="282">Largusoperla</emphasis>
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. It can be easily distinguished from all other congeners by the paraproct strongly sclerotized and subapically constricted to a needle-like spine (
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; 
<figureCitation id="6BA52A54FF9F0B241D23FC4338ECFC3F" box="[1332,1436,997,1022]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="6.[151,250,1843,1865]" captionStartId-1="7.[151,250,1802,1824]" captionStartId-2="9.[151,250,1455,1477]" captionTargetBox-0="[150,1409,849,1803]" captionTargetBox-1="[168,1406,710,1755]" captionTargetBox-2="[174,1413,699,1412]" captionTargetId-0="figure@6.[150,1436,832,1821]" captionTargetId-1="figure@7.[150,1436,661,1780]" captionTargetId-2="figure@9.[150,1436,663,1433]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="9" captionText-0="FIGURE 5. Largusoperla reni, sp. nov. Photograph and reconstruction of terminalia, ventral view." captionText-1="FIGURE 6. Available head patterns of Largusoperla, dorsal view." captionText-2="FIGURE 7. Available hammer shapes of Largusoperla, ventral view." httpUri-0="https://zenodo.org/record/2589806/files/figure.png" httpUri-1="https://zenodo.org/record/2589808/files/figure.png" httpUri-2="https://zenodo.org/record/2589810/files/figure.png" pageId="1" pageNumber="282">
Figs. 
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in Chen 
<emphasis id="C1EAEAC3FF9F0B2418E1FBAC3C5EFBE0" box="[246,302,1033,1057]" italics="true" pageId="1" pageNumber="282">et al.</emphasis>
2018).
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