Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia
Author
Jirapatrasilp, Parin
https://orcid.org/0000-0002-5591-6724
Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand
Author
Sutcharit, Chirasak
Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand
jirasak4@yahoo.com
Author
Panha, Somsak
Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand
somsak.pan@chula.ac.th
text
ZooKeys
2022
2022-08-25
1119
1
115
http://dx.doi.org/10.3897/zookeys.1119.85400
journal article
http://dx.doi.org/10.3897/zookeys.1119.85400
1313-2970-1119-1
A3BE91C6B79344E1A886A803BF104D8B
F158A7C5261D52B69288A62F3C777CAF
Family
Pupinidae Pfeiffer, 1853
Remarks.
Currently, there are three subfamilies within the family
Pupinidae
:
Pupininae
,
Liareinae
Powell, 1946, and
Pupinellinae
Kobelt, 1902 (
Bouchet et al. 2017
). The subfamily
Liareinae
was endemic to New Zealand, originally established as a family (
Powell 1946
), and this familial assignment was adopted by
Egorov (2013)
. Later,
Ponder and
Waren
(1988)
treated this taxon as a subfamily of the
Pupinidae
; this classification scheme was adopted by
Bouchet et al. (2017)
and
MolluscaBase (2022)
.
The subfamily
Pupinellinae
was originally established as a section under the
Pupinidae
, and the only diagnostic character that distinguished this subfamily from the
Pupininae
is the shell surface (
Kobelt 1902
). The
Pupininae
has a shell surface covered by glaze, which is glossy and completely smooth, whereas the shell surface of the
Pupinellinae
is without glaze, being either striated, matt or silky-shiny (
Kobelt 1902
;
Egorov 2013
). Whether this character is a subfamilial synapomorphy needs further confirmation because at least one
Pupina
species has a matt surface (e.g.,
P. arula
) and some
Pupinella
species have a somewhat glossy surface (e.g.,
P. mansuyi
,
P. illustris
).