Taxonomic revision and systematic notes on some Halecium species (Cnidaria, Hydrozoa)
Author
Schuchert, Peter
text
Journal of Natural History
2005
2005-02-28
39
8
607
639
http://dx.doi.org/10.1080/00222930400001319
journal article
10.1080/00222930400001319
1464-5262
4668977
Halecium petrosum
Stechow, 1919
(
Figure 8
)
Halecium robustum
:
Motz-Kossowska 1911
, p 346
,
Figure 14
;
Neppi 1921
, p 13,
Figures 4
,
5
.
[Not
Halecium robustum
Pieper 1884
, p 166
.]
Halecium petrosum
Stechow 1919
, p 36
, new name;
Coma et al. 1992
, p 162,
Figure 1A
;
Peña Cantero and García Carrascosa 2002
, p 69,
Figure 14
a–c.
Figure 8.
Halecium petrosum
Stechow, 1919
; after material from Banyuls-sur-Mer. (A) Colony silhouette (scale bar: 5 mm); (B) two internodes (scale bar 0.2 mm); (C) branching point (same scale bar as B); (D) hydrophore, primary and secondary hydrothecae (scale bar: 0.1 mm); (E) hydrotheca with straight walls (same scale bar as D); (F) female gonotheca in side view (same scale bar as B); (G) hydrotheca of female gonotheca in anterior view (anterior side in F is directed towards left) (same scale bar as D); (H) part of male colony, sperm masses stippled, note that this sample shows signs of multiple re-growth and regeneration, its identity is not entirely secure (scale bar: 0.2 mm).
Material examined
MHNG
INVE 26665
,
Anse de Troc
,
Banyuls-sur-Mer Pyrénées Orientales
,
France
,
2 m
, coll.
P. Schuchert
,
23 June 1997
, fertile female
.
MHNG
INVE 29763
,
Spain
,
Mallorca
,
Cala Murada
, coll.
P. Schuchert
,
23 August 2000
,
0.5 m
, on rock and red algae, shows signs of multiple re-growth and regenerations, identification not entirely secure
.
Description
Colonies erect, branched, shoots
5–15 mm
, monosiphonic, composed of main axis and shorter side-branches. Erect parts segmented by alternately oblique nodes; nodes distinct, internodes about
0.5–0.6 mm
long, diameter
80–90 mm
. Perisarc smooth. Side-branches originate from hydrophores. Hydrotheca near segment end, on very short hydrophore; hydrotheca does not reach beyond level of node. Primary hydrotheca with straight or slightly curved walls, basal diameter
0.10–0.12 mm
, depth
20–30 mm
, desmocytes present. Hydrophore without pseudodiaphragm. Secondary hydrotheca on corrugated hydrophore of variable length, several times longer than primary hydrophore. Gonothecae on main stem. Female gonotheca
0.6–0.7 mm
long and about
0.5 mm
broad, main body ovoid, bilaterally symmetric through a lateral hydrotheca that overtops gonotheca body, with crease-line running from hydrotheca along gonotheca body. Male gonothecae without hydrotheca, club-shaped, slightly curved or not, length
0.5–0.55 mm
.
Additional information
Stems may reach
3 cm
in height. Female gonotheca with two hydranths (
Motz-Kossowska 1911
).
Biology
Depth range
1–200 m
, grows on a variety of solid substrata. Fertile colonies observed from June to October (
Peña Cantero and García Carrascosa 2002
). Ecological aspects were investigated by
Coma et al. (1992)
.
Distribution
Mediterranean.
Type
localities:
Cap Béar
and
Cap Abeille
near
Banyuls-sur-Mer
,
France
,
Mediterranean
.
Remarks
Motz-Kossowska (1911)
identified abundant material from Banyuls as
Halecium robustum
Pieper, 1884
. The latter name was recognized by
Bedot (1912
, p 14) as a synonym of
H. lankesterii
(see also
Hamond 1957
for more invalid homonyms of
Halecium robustum
). Because Motz-Kossowska’s material had female gonothecae that differed from typical
H. lankesterii
,
Stechow (1919)
attributed it to a new species with the name
Halecium petrosum
(compare
Figures 8F
and
7E
). Also the trophosomes of
Halecium lankesterii
and
H. petrosum
are quite distinct (compare
Figures 7B
and
8B
, and remarks under
H. lankesterii
).