Taxonomy, annotated new records and a checklist of Mymaridae (Hymenoptera) of Finland, with description of a new species of Eustochus
Author
Triapitsyn, Serguei V.
Entomology Research Museum, Department of Entomology, University of California, Riverside, California, 92521, USA;
Author
Koponen, Martti
Tuoppitie
Author
Vikberg, Veli
Liinalammintie 11 as.
Author
Várkonyi, Gergely
Finnish Environment Institute SYKE, Biodiversity Centre, FI- 88900, Kuhmo, Finland; e-mail: gergely. varkonyi @ ymparisto. fi
text
Acta Entomologica Musei Nationalis Pragae
2020
2020-11-06
60
2
565
589
http://dx.doi.org/10.37520/aemnp.2020.39
journal article
10.37520/aemnp.2020.39
1804-6487
5177724
659934F0-6B73-4571-8ADA-5E14CD3669EC
Erythmelus
(
Erythmelus
)
agilis
(Enock, 1909)
are accepted here and expanded as follows: clava of fe-
Material examined.
Ta
:
Janakkala, Kalpalinna (6757:8369),
7.vii.2017
, male antenna 1- (entire), 2- or 3-segmented.
Caraphractus
V. Vikberg
(
1 ♀
,
VVCT
). Walker, 1846 was considered to be most closely related to
Eustochus
of which the former might be a derived offshoot;
Distribution.
Palaearctic (
Belgium
,
Bulgaria
, Czech their only more or less significant difference was the claval Republic,
Denmark
,
Finland
,
Germany
,
Greece
,
Italy
, segmentation of the female antenna: entire in the former but
Kyrgyzstan
,
Netherlands
, North
Macedonia
,
Norway
, 2- or 3-segmented in the latter genus (Hඎൻൾ*© & Bൺඊඎൾ*©ඈ
Russia
,
Serbia
,
Sweden
,
Switzerland
,
United Kingdom
) 2007). Both genera were known from eggs of Coleoptera and Nearctic (
Canada
,
USA
) Regions (T*©ංൺඉංඍඌඒඇ 2003b; (T*©ංൺඉංඍඌඒඇ 2012; P*©ං*öඈඉ & MඈǤඅൺඇ 2016) and are T*©ංൺඉංඍඌඒඇ et al. 2007; Sൺආκඈඏග et al. 2020). strictly Holarctic in distribution (Hඎൻൾ*© & Bൺඊඎൾ*©ඈ 2007).
However, our discovery of two new species of
Eustochus
Eustochus
Haliday, 1833
from
Finland
and
Germany
with an entire clava (see be-
Caraphractus
Walker, 1846
,
syn. nov.
, downgraded to a subgenus of low) shows that the variable segmentation of the clava in
Eustochus
is not a good genus-defining character, hence
Remarks.
The diagnoses of
Eustochus
Haliday, 1833
by the proposed synonymy of
Caraphractus
syn. nov.
with Hඎൻൾ*© & Bൺඊඎൾ*©ඈ (2007) and P*©ං*öඈඉ & MඈǤඅൺඇ (2016)the earlier described
Eustochus
; furthermore, the former is treated here as its subgenus,
E.
(
Caraphractus
) stat. revid.
Caraphractus
, with its sole valid member, the
type
species
C. cinctus
Walker, 1846
(=
Eustochus
(
Caraphractus
)
cinctus
(Walker, 1846)
,
comb. nov.
), is very likely essentially nothing more than just an aquatic species of
Eustochus
. However, while proposing this new generic synonymy, we also downgrade the taxonomic status of
Caraphractus
to that of a subgenus of
Eustochus
for the following reasons kindly provided by John T. Huber (personal communication): in female
Caraphractus
, mandible with 3 serrated teeth (2, not serrated teeth in most
Eustochus
s. str.
except with 3 teeth in
E.
(
Eustochus
)
koponeni
); petiole ventrally with strong reticulations (smooth in most
Eustochus
s. str.
except in
E.
(
Eustochus
)
koponeni
) and without a longitudinal suture (suture present at least partially in most
Eustochus
s. str.
except in
E.
(
Eustochus
)
koponeni
); first gastral tergum laterally with about 10 longish setae (such setae absent in
Eustochus
s. str.
); mps absent on funicle segments (present on several segments in
Eustochus
s. str.
). Also, male
Caraphractus
have 1 less antennal flagellar segment than male
Eustochus
s. str.
, but these are known only for
E.
(
Eustochus
)
atripennis
(Curtis, 1832)
.