A revised classification of the genus Matrona Selys, 1853 using molecular and morphological methods (Odonata: Calopterygidae) Author Yu, Xin Author Xue, Junli Author Hämäläinen, Matti Author Liu, Yang Author Bu, Wenjun text Zoological Journal of the Linnean Society 2015 2015-06-23 174 3 473 486 http://dx.doi.org/10.1111/zoj.12253 journal article 10.1111/zoj.12253 0024-4082 10107117 SUBGENUS MATRONA SELYS, 1853 Generic diagnosis: Male wings uniformly dark blue opaque or reddish brown with a distinctive area of the wing base, of variable size, which appears milky when viewed from an oblique angle, owing to the presence of bluish white cross-veins. The reticulation of the wings is dense to very dense, especially in the hind wings ( Fig. 6A, B ). Median space usually with 5–7 reticulated cross-veins.