New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae) Author Perkins, Philip D. text Zootaxa 2011 2011-10-28 3074 1 198 journal article 31530 10.5281/zenodo.1050060 2d28bebf-e3c5-457b-9152-8a6877ded281 1175-5326 1050060 Steineri Complex Two species comprise the Steineri Complex: H. steineri , and H. concepcionica . These are medium sized species (ca. 1.49–1.52 mm ), which have coarse and dense dorsal punctation, lack a pronotal scintilla, and have similarly shaped metaventral plaques ( Figs. 118–119 ). The male genitalia in this complex are comparatively simplified, rather wide in lateral aspect, the main piece with a more or less developed, raised ridge on the ventral surface, and the gonopore is located at the end of a long flexible flagellum ( Figs. 120–121 ). The currently known species are from Bolivia ( H. steineri ) and Paraguay ( H. concepcionica ).