Revision of the genus Orictites ANDREWES, 1931 (Coleoptera, Carabidae, Clivinini) Author Balkenohl, Michael text Contributions to Natural History 2017 2017-07-17 35 1 68 journal article 20497 10.5169/seals-787048 8c468ca9-0ee3-4200-b5a3-6adfd98caaea 2624-9170 6284316 Orictites costulipennis BATES, 1892 , comb. nov. Catalogue Clivina costulipennis ; Bates 1892: 281 ; Clivina costulipennis BATES ; Andrewes 1926: 373 ; Clivina costulipennis BATES ; Andrewes 1929: 366 ; Clivina costulipennis BATES ; Andrewes 1930: 113 ; Clivina costulipennis BATES ; Lorenz 1998: 131 ; Clivina costulipennis BATES ; Balkenohl 2001: 14 . Type material: Holotype : ♂ , labels: brown with black frame, black printed: “ Palon ( Pegú ) L.Fea.VIII. IX.87 ” / brown, handwritten in black: “ Clivina costulipennis Bates ” / yellowish-brown, handwritten in black: “ Cliv. costulipennis (typus!) Bates” / white, red-printed with red frame: “TYPUS” / brown, printed in black: “Mus. Civ. Genova” / white with black frame, handwritten in black: “ costulipennis Bates ” / red with black frame, black printed “HOLOTYPUS” and handwritten “ Clivina costulipennis Bates, 1892 ” Additional label: white, black printed: “Genitalpräparat wasserlÖslich M. Balkenohl I 07 ” ( MCSNG ) . Additional material: 2 ♂ , 1 ♀ , 3 specimens : Thailand , Doi Inthanon , 4.–6.5.1990 , 8.–9.5.1990 , 23.–25.5.1990 , 12.–19.6.1990 , leg. Malicky / Clivina costulipennis BAT. det. M.Baehr’ 94 ( CBM-ZSM , CBB ); 1 specimen : Penang ,G.E.Bryant. 13.XI.13 / H.E. Andrewes Coll. B.M.1945–97. / Clivina costulipennis BATES Compared with type H.E.A. ( BMNH ); 1 specimen : Malay Pa. Perak / 58J81 / Doherty / Fry Coll 1905.100. / Clivina costulipennis BATES Compared with type H.E.A.; 1 specimen : NE-INDIA: Meghalaya W Garo Hills Norek NP; ca. 1100 m / 25°25.6´N 90°19.5´E 9.–17.5.1996 leg. Jendek & Sausa ( NHMW ); 2 ♂ , 3 ♀ , W. THAILAND : 300m ., Thung Yai Wildlife Sanctuary. 15°28´N – 98°48´E . / Tak Province Umphang District Song Bae Stream . 18–27.iv.1988 . / Evergreen rain forest M.J.D.Brendell. B.M.1988–183. ( BMNH , CBB ); 1 ♂ , Thailand , 23.–25.ii.1996 Ranong prov. 9°57´98°45´ Hat Som Paen env . K. Majer leg. ( CBP ); 1 ♀ , NE INDIA Meghalaya Nokrek n.p.,3kmSDaribokgirl 25°27´N , 90°19´E , 26.4.1999 Dembický & Pacholátko lgt. ( CBB ); 1 specimen : N. THAILAND , Chiang Mai prov. San Pakia , 19°19’N , 98°50’ E , 1400 m , 1.–15.5.1998 , Michal Bednařik leg. ( CBP ); 1 specimen : NE INDIA , MEGHALAYA , SW of CHERRAPUNJEE 25°13´–15´N 91°40´E; 900m L. Dembický leg.; 11.– 12.v.2004 ( NHMB ). Redescription ( Figs 2 , 16 , 18 , 20 , 32 ; pp. 8, 15, 19, 23, 35) Measurements: Holotype : Body length 5.1 mm , width 1.45 mm ; ratio length/ width of pronotum 0.98; ratio length/width of elytra 1.91. Additional material (n = 10): Body length 5.5–6.5 mm (  = 5.6 mm ), width 1.49–1.75 mm (  = 1.62 mm ), ratio length/width of pronotum 0.93 – 1.02 (  = 0.96), ratio length/width of elytra 1.8 – 1.95 (  = 1.87). Colour: Shiny. Black to dark-brown. Wings of clypeus and supraantennal plates more or less transparent red-brown, antennae, labrum and palpi yellow brown, legs red-brown. Head: A fifth narrower than pronotum. Clypeus with small rounded tooth laterally, anterior margin slightly excised, with raised knob between the two clypeal setae, posterior part of knob covered with big and medium sized punctures; margin of wings convex, hollowed out, divided from middle part of clypeus by slight notches and carinae, divided from supraantennal plates by distinct obtusely angled notches and ridge. Clypeus and wings reflexed margined, supraantennal plates vaulted up to mid-eye level, smooth, with reflexed rounded margin, elongated as supraorbital carina, frons separated from clypeus by flat transverse depression and group of big punctures, with costate, nearly parallel running frontal carinae at each side posteriorly, with broad furrow between frontal carina and supraorbital carina, with flat punctured transverse furrow, sharp furrow between frons and frontal carina. Frons moderately convex, with very minute scattered punctures. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from tubercle. Eyes of moderate size, flattened but still convex, genae enlarged, regularly rounded, not as high as and shorter than eyes, enclosing eyes posteriorly, forming distinct angle at neck. Grooves for reception of antennae short, length 0.4 of eye diameter. Neck constriction marked as distinct step, with band of bigger punctures almost doubled, neck laterally covered with reticulation and densely scattered medium sized punctures. Labrum straight, 7-setose, ciliate laterally, with isodiametric reticulation. Mandibles robust, nearly as long as head, flattened, stout, broadened at base, acutely curved at apex, carinae of scrobe complete, both mandibles obtusely angled towards base (opened position), both mandibles somewhat hollowed out dorsally. Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, slightly longer than 2 nd segment, 2 nd segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck covered with bigger sized punctures and transverse reticulation, Submentum and mentum distinctly separated; submentum with four bigger, about equally spaced setigerous punctures; mentum at base bilaterally with broad and rounded tubercle, with some longitudinal carinae, with fine and complete reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth forming a broad obtuse triangle, bisetose, not projecting as far as lateral lobes. Paragenae sharp, carina-like, with distinct tooth anteriorly, rounded at base. Antennae short, reaching up to middle of pronotum, scapus knee-like angled, with one seta dorsoapically, scapus and pedicellus with fine reticulation, segments 5–10 moniliform, antennomeres somewhat flattened, with shiny areas on flattened parts, segments 4–11 densely pubescent, with few additional long setae. Pronotum: Square, as long as wide, sides slightly convex at middle, distinctly rounded in anterior quarter to anterior angles. Anterior margin straight. Reflexed lateral margin smooth, from posterior angle to base running as a straight line. Lateral channel conspicuously broad between the two lateral setigerous punctures, with isodiametric reticulation and very few irregular punctures. Anterior setigerous puncture located at the end of anterior quarter approximated to the convexity of the disc of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angle projecting, rounded, formed by reflexed lateral margin, posterior angle developed as sharp distinct tooth, projecting laterally. Anterior transverse line formed by distinct, partly connected punctures. Median line deep and conspicuously broad, running up to level of anterior transverse line but not joining, smaller at base, adjoining base. Surface scattered with some fine to medium sized punctures and with isodiametric reticulation laterally and at base, basal impression indistinct, basal channel broad, deep. Flange raised keel-like (lateral view). Elytra: Subcylindrical, sides slightly diverging, marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles. Reflexed margin with fine crenulation in anterior quarter, margin slightly thickened at humerus, bending with channel over rounded humerus up to 5 th stria. No humeral tooth. Basal tubercle distinct, with setigerous puncture, situated at declivity of 1 st stria. Striae 1–4 free at base, all striae deep, broad, more or less distinctly punctuate, striae 1–2, 3–4 and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura costiform, raised. All intervals conspicuously convex, convexity increasing laterally, thought costate, 3 rd and 4 th broader and raised at base, both with tubercle at base, 8 th carinate. Interval 3 with four setigerous punctures adjoining stria 3. Hind wings: Fully developed. Ventral surface: Epipleuron shiny, covered with medium sized scattered punctures and indistinct isodiametric reticulation, submarginal furrow distinct. Proepisternum and episternum nearly smooth, prosternite smooth, keeled at middle like an inverted V, mesosternum smooth. Sternites with transverse to isodiametric reticulation, each with big punctures at base laterally, 3 rd to 5 th with paralateral ambulatory setae at each side, ventral strigae distinct, sternite six smooth at middle, with two widely separated apical setae at each side. Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral, no dorsal furrow, basal tarsomere elongated, nearly as long as the following four together. Mesotibia with preapical spine, furnished with strong seta. All tarsomeres relatively broad. Male genitalia ( Fig. 20 ): Relatively long, moderately sclerotized. Median lobe slender, moderately arcuate, slightly twisted apically. Endophallus strongly folded, teeth not visible. Parameres of about same length, both slender, slightly twisted, asetose. Female genitalia ( Fig. 32 ): Coxostylus slightly sigmoid, dorso-ventrally moderately depressed, gently curved to apex, with one seta at apex, with one longer ensiform and four nematiform setae in middle part. Variation: The anterior transverse line of the pronotum is more or less deeply engraved. The isodiametric reticulation in the lateral channel of the pronotum is more or less distinctly developed. Specimens from Sabah and Sarawak ( Malaysia ) have the tubercles on the 3 rd and 4 th interval at base indistinctly developed. The number of punctures on the disc of the pronotum varies. Diagnosis: A medium-sized almost black species with distinct knob on the clypeus and distinct flat transverse punctured depression behind, without humeral tooth, tubercles at the base of intervals 3 and 4 of the elytra, and four setigerous punctures on interval 3. Distinguished from all other species by the combination of the following characters: The frons of the head is covered by minute scattered punctures, the reflexed lateral margin of the pronotum is smooth, and all intervals of the elytra are conspicuously convex. Distribution: Known from North, middle and Western parts of Thailand and from Megalaya (NE India ). Some specimens were collected at 450 m up to 1400 m in litter of forests. Remarks: Bates based the description on one male specimens deposited in MCSNG and he described the species as “curious” ( Bates 1892: 282 ) obviously due to the striking differences to the Clivina species described in the same contribution. Kult (1951) did not mention the species at all.