<document id="9038D23032F4F64A72BF9E2ACBE85834" ID-CLB-Dataset="7439" ID-DOI="10.11646/zootaxa.4934.1.1" ID-GBIF-Dataset="300bac18-55f5-4e34-90e1-7fccd02d68c1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4558796" ID-ZooBank="79895443-4597-42A5-AF8A-023EACB20E10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614157470936" checkinUser="plazi" docAuthor="Archibald, Bruce, Cannings, Robert A., Erickson, Robert J., Bybee, Seth M. &amp; Mathewes, Rolf W." docDate="2021" docId="03B487C2007FFFF9FF5BF923FD941308" docLanguage="en" docName="zootaxa.4934.1.1.pdf" docOrigin="Zootaxa 4934 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Whetwhetaksa Archibald &amp; Cannings 2021, new genus" docType="treatment" docVersion="6" lastPageNumber="111" masterDocId="FF8DFFBA0012FF97FFCCFFE5FFAE1215" masterDocTitle="The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America" masterLastPageNumber="133" masterPageNumber="1" pageNumber="110" updateTime="1698931040708" updateUser="ExternalLinkService">
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<mods:title id="D1A0A720E39516CC936217729DF5AF77">The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America</mods:title>
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<mods:namePart id="564A72E5FAD90C7C302904EE4B18A3F9">Archibald, Bruce</mods:namePart>
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<mods:namePart id="2F0BB03E8DBD4B124CA95BD6C2CF4556">Cannings, Robert A.</mods:namePart>
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<mods:namePart id="5189DCA7E49AB994759678AA0EB0665B">Erickson, Robert J.</mods:namePart>
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<mods:namePart id="76477B66E60BC880ACAB96FFCB9F8428">Mathewes, Rolf W.</mods:namePart>
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<treatment id="03B487C2007FFFF9FF5BF923FD941308" ID-DOI="http://doi.org/10.5281/zenodo.4558866" ID-GBIF-Taxon="180881785" ID-Zenodo-Dep="4558866" LSID="urn:lsid:plazi:treatment:03B487C2007FFFF9FF5BF923FD941308" httpUri="http://treatment.plazi.org/id/03B487C2007FFFF9FF5BF923FD941308" lastPageId="110" lastPageNumber="111" pageId="109" pageNumber="110">
<subSubSection id="C307655F007FFFFAFF5BF923FD5414F4" box="[151,762,1734,1761]" pageId="109" pageNumber="110" type="nomenclature">
<paragraph id="8BA236D4007FFFFAFF5BF923FD5414F4" blockId="109.[151,762,1734,1797]" box="[151,762,1734,1761]" pageId="109" pageNumber="110">
<heading id="D0EA81B8007FFFFAFF5BF923FD5414F4" bold="true" box="[151,762,1734,1761]" fontSize="11" level="2" pageId="109" pageNumber="110" reason="4">
<emphasis id="B969EAC6007FFFFAFF5BF923FD5414F4" bold="true" box="[151,762,1734,1761]" pageId="109" pageNumber="110">
<taxonomicName id="4C1D4D57007FFFFAFF5BF923FDC214F5" authority="Archibald &amp; Cannings" authorityName="Archibald &amp; Cannings" authorityYear="2021" box="[151,620,1734,1761]" class="Insecta" family="Sieblosiidae" genus="Whetwhetaksa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B969EAC6007FFFFAFF5BF923FEE914F4" bold="true" box="[151,327,1734,1761]" italics="true" pageId="109" pageNumber="110">Whetwhetaksa</emphasis>
Archibald &amp; Cannings
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, 
<taxonomicNameLabel id="A25A57BD007FFFFAFDB4F922FD5414F4" box="[632,762,1735,1761]" pageId="109" pageNumber="110" rank="genus">new genus</taxonomicNameLabel>
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</heading>
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<paragraph id="8BA236D4007FFFFAFF5BF90EFE8D1510" blockId="109.[151,762,1734,1797]" box="[151,291,1771,1797]" pageId="109" pageNumber="110">
<figureCitation id="13262A51007FFFFAFF5BF90EFEB11510" box="[151,287,1771,1797]" captionStart-0="FIGURE 66" captionStart-1="FIGURE 67" captionStart-2="FIGURE 68" captionStart-3="FIGURE 69" captionStart-4="FIGURE 70" captionStartId-0="112.[151,250,1985,2009]" captionStartId-1="113.[151,250,1984,2008]" captionStartId-2="114.[151,250,1961,1985]" captionStartId-3="115.[151,250,1805,1829]" captionStartId-4="116.[151,250,1955,1979]" captionTargetBox-0="[343,1248,184,1961]" captionTargetBox-1="[269,1319,187,1957]" captionTargetBox-2="[399,1189,182,1937]" captionTargetBox-3="[151,1423,181,1769]" captionTargetBox-4="[272,1316,181,1931]" captionTargetId-0="figure-19@112.[326,1261,181,1961]" captionTargetId-1="figure-17@113.[257,1330,181,1960]" captionTargetId-2="figure-19@114.[386,1202,181,1937]" captionTargetId-3="figure-17@115.[151,1436,181,1781]" captionTargetId-4="figure-19@116.[257,1331,181,1931]" captionTargetPageId-0="112" captionTargetPageId-1="113" captionTargetPageId-2="114" captionTargetPageId-3="115" captionTargetPageId-4="116" captionText-0="FIGURE 66. Whetwhetaksa millerae holotype, SR 06-01-38. A, photograph; B, drawing. Both to scale, 5 mm." captionText-1="FIGURE 67. Whetwhetaksa millerae paratype 1, SR 10-36-10. A, photograph; B, drawing. Both to scale." captionText-2="FIGURE 68. Whetwhetaksa millerae paratype 2, SRUI 99-98-34. A, photograph; B, drawing. Both to scale; aa, accessory antenodal crossvein." captionText-3="FIGURE 69. Whetwhetaksa millerae additional material. SR 93-11-01: A, photograph; B, drawing; SR 06-01-36: C, photograph; D, drawing.All to scale, 5 mm." captionText-4="FIGURE 70. Whetwhetaksa millerae, additional material. SR 10-14-19: A, photograph; B, drawing; SR 05-11-01: C, photograph; D, drawing.All to scale, 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4558960" figureDoi-1="http://doi.org/10.5281/zenodo.4558964" figureDoi-2="http://doi.org/10.5281/zenodo.4558968" figureDoi-3="http://doi.org/10.5281/zenodo.4558970" figureDoi-4="http://doi.org/10.5281/zenodo.4558972" httpUri-0="https://zenodo.org/record/4558960/files/figure.png" httpUri-1="https://zenodo.org/record/4558964/files/figure.png" httpUri-2="https://zenodo.org/record/4558968/files/figure.png" httpUri-3="https://zenodo.org/record/4558970/files/figure.png" httpUri-4="https://zenodo.org/record/4558972/files/figure.png" pageId="109" pageNumber="110">Figs. 66–70</figureCitation>
.
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<emphasis id="B969EAC6007FFFFAFF5BF8D7FEBC1559" bold="true" box="[151,274,1842,1868]" pageId="109" pageNumber="110">Diagnosis.</emphasis>
As for family.
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<subSubSection id="C307655F007FFFFAFF0BF8B2FDD81564" box="[199,630,1878,1905]" pageId="109" pageNumber="110" type="type_taxon">
<paragraph id="8BA236D4007FFFFAFF0BF8B2FDD81564" blockId="109.[151,1437,1842,2013]" box="[199,630,1878,1905]" pageId="109" pageNumber="110">
<emphasis id="B969EAC6007FFFFAFF0BF8B2FECC1565" bold="true" box="[199,354,1878,1905]" pageId="109" pageNumber="110">
<typeStatus id="54A68876007FFFFAFF0BF8B2FEAC1564" box="[199,258,1879,1905]" pageId="109" pageNumber="110">Type</typeStatus>
species.
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<taxonomicName id="4C1D4D57007FFFFAFEA5F8B2FDC11564" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[361,623,1879,1905]" class="Insecta" family="Sieblosiidae" genus="Whetwhetaksa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="millerae">
<emphasis id="B969EAC6007FFFFAFEA5F8B2FDC11564" box="[361,623,1879,1905]" italics="true" pageId="109" pageNumber="110">Whetwhetaksa millerae</emphasis>
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.
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<subSubSection id="C307655F007FFFFAFF0BF89FFC351580" box="[199,923,1914,1941]" pageId="109" pageNumber="110" type="description">
<paragraph id="8BA236D4007FFFFAFF0BF89FFC351580" blockId="109.[151,1437,1842,2013]" box="[199,923,1914,1941]" pageId="109" pageNumber="110">
<emphasis id="B969EAC6007FFFFAFF0BF89FFEF71581" bold="true" box="[199,345,1914,1940]" pageId="109" pageNumber="110">Description.</emphasis>
With the character states of its only species, below.
</paragraph>
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<subSubSection id="C307655F007FFFF9FF0BF87AFD941308" lastPageId="110" lastPageNumber="111" pageId="109" pageNumber="110" type="etymology">
<paragraph id="8BA236D4007FFFF9FF0BF87AFECC12C0" blockId="109.[151,1437,1842,2013]" lastBlockId="110.[151,1437,151,285]" lastPageId="110" lastPageNumber="111" pageId="109" pageNumber="110">
<emphasis id="B969EAC6007FFFFAFF0BF87AFEE615AC" bold="true" box="[199,328,1951,1977]" pageId="109" pageNumber="110">Etymology</emphasis>
. The genus name is formed from the word in the language of the Colville Indian tribe x̌ʷətx̌ʷətaqs, meaning odonate, pronounced “ 
<emphasis id="B969EAC6007FFFFAFE32F826FD2315C8" box="[510,653,1987,2013]" italics="true" pageId="109" pageNumber="110">whetwhetaks</emphasis>
”. The Republic site where the only known fossils of this family have been found is on the traditional land of the Colville Indian Tribe, immediately north of the current Colville Reserve. Gender, feminine.
</paragraph>
<paragraph id="8BA236D4007CFFF9FF0BFF3AFD941308" blockId="110.[151,1437,151,285]" pageId="110" pageNumber="111">
<emphasis id="B969EAC6007CFFF9FF0BFF3AFED312EC" bold="true" box="[199,381,223,249]" pageId="110" pageNumber="111">Range and age</emphasis>
. Republic localities B4131 and A0307B of the Tom Thumb Tuff Member of the Klondike Mountain Formation; latest Ypresian.
</paragraph>
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</treatment>
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