Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species
Author
Fallon, Phillip J.
text
Zootaxa
2016
4090
1
1
363
journal article
10.11646/zootaxa.4090.1.1
e6b35f5a-435e-4473-b29e-1e4d842f84b0
1175-5326
263299
203BAC25-B542-48FE-B5AD-EBA8C0285833
Calliclava fasciata
,
new species
(
Plate 17
)
Type
material.
Holotype
, 16.1 x 6.0 mm (UF 470272); 2
paratypes
: 1 spec., 17.3 x
6.6 mm
, in
6 m
, Los Roques,
Venezuela
(USNM 1291323); 1 spec., 17.6 x
6.8 mm
, in
22–25 m
, off Cabo Codera, Miranda State,
Venezuela
(P. Stahlschmidt coll.).
Type
locality.
Arenas Point,
Tortuga
I.,
Venezuela
, in
8 m
.
Range and habitat.
Venezuela
(
Tortuga
I.; Los Roques Is.). If the specimen depicted in Daccarett & Bossio (2011, species 787) is this species, then its range would include
Colombia
. Specimens from Venezuelan localities were taken from
6–25 m
depths. The Colombian specimen, if proven to be this species, was reported from
44 m
.
Description.
Shell
small (to
17.6 mm
total length), fusiform, glossy, translucent between ribs and spirals; whorls convex, about 10½ total, the last approximately 56% of total length; sutures impressed; principal sculptural element of ribs and spiral threads. Aperture narrow, widest posteriorly near anal sinus.
Protoconch
approximately 2¼–2½ translucent whorls, a clearly visible carina present on both whorls beginning mid-whorl on the first, subperipheral beginning on 2nd to end of protoconch.
Axial sculpture
of numerous narrow, opisthocline ribs, that extend from suture-to-suture, are recurved in sulcus, and run to anterior fasciole on last whorl; whorls number 14– 16 on penultimate, 9–12 on last whorl to varix; rib interspaces much wider than ribs. Growth striae present over entire shell, most visible between spiral threads.
Varix
broad, hump-like, approximately ⅓-turn from edge of outer lip.
Spiral sculpture
of flattened threads, interspaces less than thread width, or alternatively, wide grooves evenly spaced creating the appearance of flattened threads, closer and narrower in sulcus; threads override axials giving them a somewhat nodulose appearance; threads number 6 on penultimate and 15 on last whorl to anterior fasciole. An additional 7 present on fasciole.
Sulcus
convex, about ¼- to ⅓-whorl height with 3 spiral threads in addition to axial ribs that are narrowed, educed, and recurved.
Outer lip
flattened from varix to edge of lip thus projecting out from the circumference of the shell in apical view; with 4 axial folds; lip edge forms a flat arc from anal sinus to the stromboid notch. Stromboid notch deep, edge flared.
Anal sinus
deep, positioned whorl shoulder, U-shaped, entrance slightly constricted by callus on parietal wall; upward flare of sinus edge and outward curve of outer lip gives the appearance of being spout-like.
Inner lip
erect anteriorly, recumbent and wider on parietal wall, and formed into a lobe posteriorly; incremental growth layers visible at edge of lip.
Anterior canal
open, narrow, moderately long, turned to the right when viewed ventrally, upturned slightly at its tip, and notched. Anterior fasciole not swollen.
Color
white with a wide yellow-brown band mid-whorl, which appear on the bottom half of spire whorls, and darker brown patch on varix. Aperture light violet, which shows through the outer lip.
Remarks.
Taxonomy.
Calliclava fasciata
has all of the characteristics of the genus: numerous ribs that extend from suture-to-suture with a distinctly demarcated sulcus, conspicuous spiral threads across intercostal spaces, and a protoconch with both whorls carinate.
Variability.
The
3 specimens
have an average total length of 17.00 mm (
16.1–17.6 mm
) and an average W/
L ratio
of 0.377.
Identification.
Calliclava fasciata
is similar to
C. vigorata
,
new species
. It is somewhat larger (17.00 versus
13.11 mm
average total length), has a yellow-brown mid-whorl band (
C. vigorata
is usually white), and not as stout (average W/L = 0.377 versus 0.399). From
C. tobagoensis
,
new species
it differs in size, coloration, and
minor
details of surface sculpture.
Calliclava fasciata
is larger (17.0 versus
10.8 mm
average total length), and has a white shell with a wide, mid-whorl spiral band of color;
C. tobagoensis
has a more uniformly colored shell with maculation in the intercostal spaces on the whorl shoulders.
Etymology.
The Banded
Calliclava
; named for the prominent yellow-brown mid-whorl spiral band present on all specimens examined. From the Latin adjective
fasciata
meaning having a band.