Revision of the genus Fascaplysinopsis, the type species Fascaplysinopsis reticulata (Hentschel, 1912) (Porifera, Dictyoceratida, Thorectidae) and descriptions of two new genera and seven new species
Author
Ekins, Merrick
Queensland Museum, PO Box 3300, South Brisbane 4101, Brisbane, Queensland, Australia & Griffith Institute for Drug Discovery, Griffith University, Brisbane 4111, Queensland, Australia & School of Biological Sciences, University of Queensland, St Lucia, Queensland, 4072 Australia
Author
Erpenbeck, Dirk
0000-0002-6380-7421
Dept. of Earth and Environmental Sciences Ludwig-Maximilians-Universität M ¸ nchen, Richard-Wagner-Strasse 10, 80333 Munich, Germany & GeoBio-Center, Ludwig-Maximilians-Universität M ¸ nchen, Richard-Wagner-Strasse 10, 80333 Munich, Germany & SNSB-Bayerische Staatssammlung f ¸ r Paläontologie und Geologie, Richard-Wagner-Str. 10, 80333 M ¸ nchen, Germany woerheide @ lmu. de; https: // orcid. org / 0000 - 0002 - 6380 - 7421
woerheide@lmu.de
Author
Debitus, Cécile
0000-0002-3978-0644
IRD, University of Brest, CNRS, Ifremer, LEMAR, F- 29280 Plouzané, France & IRD, Univ de la Polynésie française, Ifremer, ILM, EIO, F- 98713 Papeete, French Polynesia & cecile. debitus @ ird. fr; https: // orcid. org / 0000 - 0002 - 3978 - 0644
cecile.debitus@ird.fr
Author
Petek, Sylvain
0000-0001-7920-0467
IRD, University of Brest, CNRS, Ifremer, LEMAR, F- 29280 Plouzané, France & IRD, Univ de la Polynésie française, Ifremer, ILM, EIO, F- 98713 Papeete, French Polynesia & sylvain. petek @ ird. fr; https: // orcid. org / 0000 - 0001 - 7920 - 0467
sylvain.petek@ird.fr
Author
Mai, Tepoerau
IRD, Univ de la Polynésie française, Ifremer, ILM, EIO, F- 98713 Papeete, French Polynesia & Institut Louis Malardé-Papeete, BP 30 - 98713 Papeete, Tahiti, French Polynesia
Author
Wörheide, Gert
0000-0002-6380-7421
Dept. of Earth and Environmental Sciences Ludwig-Maximilians-Universität M ¸ nchen, Richard-Wagner-Strasse 10, 80333 Munich, Germany & GeoBio-Center, Ludwig-Maximilians-Universität M ¸ nchen, Richard-Wagner-Strasse 10, 80333 Munich, Germany & SNSB-Bayerische Staatssammlung f ¸ r Paläontologie und Geologie, Richard-Wagner-Str. 10, 80333 M ¸ nchen, Germany woerheide @ lmu. de; https: // orcid. org / 0000 - 0002 - 6380 - 7421
woerheide@lmu.de
Author
Hooper, John N. A.
0000-0003-1722-5954
Queensland Museum, PO Box 3300, South Brisbane 4101, Brisbane, Queensland, Australia & john. hooper @ qm. qld. gov. au; https: // orcid. org / 0000 - 0003 - 1722 - 5954
john.hooper@qm.qld.gov.au
text
Zootaxa
2023
2023-09-19
5346
3
201
241
http://dx.doi.org/10.11646/zootaxa.5346.3.1
journal article
10.11646/zootaxa.5346.3.1
1175-5326
8390072
C577D701-4F0A-44AB-8CAF-9DF56BEEAA9C
Genus
Fascaplysinopsis
Bergquist, 1980
Definition
: Superficially armoured
Thorectinae
, with a dense collagenous matrix supported by a skeleton of thick cored heavily laminated primary fibres, uncored laminated secondary and tertiary fibres, often with pithing.
Diagnosis.
Lobate upright digitate fistular and semi burrowing sponges, with a superficial armour and rounded conules. The skeleton is semi-regular but characterised by large diameter detritus cored fasciculating laminated primary fibres, uncored often pithed, laminated secondary fibres and tertiary fibres. The primary fibres can be 2 to 10 times the diameter of the secondary fibres. The tertiary fibres can also be 2 to 10 times smaller than the secondary fibres. The mesohyl is characteristically gelatinous due to the collagenous matrix, but the consistency is firm and compressible.
Remarks.
The genus
Fascaplysinopsis
was created by
Bergquist (1980)
to accommodate irregular branched thorectid sponges with detritus-cored fasciculated primary fibres and uncored secondary fibres. This was meant to differentiate species from
Aplysinopsis
that have large fasciculated primary fibres, lack a sand cortex, and have a cavernous internal skeleton. When doing this, Bergquist designated
Aplysinopsis reticulata
Hentschel, 1912
as the type species, which had previously been synonymised with
Fasciospongia cavernosa
(
Schmidt, 1862
)
by
de Laubenfels (1948: 123)
, the latter action rejected by
Bergquist (1980)
. Unfortunately, in defining
Fascaplysinopsis
, Bergquist
relied heavily on recently collected material from
Queensland
,
Australia
. Here, we examined the
syntype
of
A. reticulata
(SMF 904) and compared it to the specimens from
Queensland
described by
Bergquist (1980)
and used in the illustration of the type species of
Fascaplysinopsis
(
Bergquist 1980
:
Fig. 16 A–B
), which is here attributed to a new species from a new genus of
Thorectinae
,
Skolosachlys
gen. nov
.
In addition, the material from
New Caledonia
and northern Queensland later added by
Bergquist (1995: 18)
to
F. reticulata
,
is also attributed here to a new species from another new genus of
Thorectinae
,
Rubrafasciculus
gen. nov.
Thus,
Fascaplysinopsis
is restricted to a monotypic genus containing only its type species,
F. reticulata
(
Hentschel, 1912
)
, but having its generic definition amended based on re-examination of the
syntype
of
A. reticulata
, here designated as the
lectotype
. The addition of new species
F. palauensis
sp. nov.
,
F. klobos
sp. nov.
and
F. ronquinni
sp. nov.
brings the total number of species in
Fascaplysinopsis
to four.