Arcofuzia cana gen. et sp. n. (Insecta, Blattaria, Fuziidae) from the Middle Jurassic sediments of Inner Mongolia, China Author Wei, Dandan Author Shih, Chungkun Author Ren, Dong text Zootaxa 2012 3597 25 32 journal article 10.5281/zenodo.283202 ede67cab-6e86-4269-9a98-f10a7df6407a 1175-5326 283202 Arcofuzia cana sp. nov. ( Figs. 1–4 ) Type specimens: Holotype male ( Figs. 1 , 2 ), CNU-BLA-NN-2011058, terminalia with forceps-like grasping cerci, a well-preserved specimen; paratype , likely male based on elongated abdominal shape, CNU-BLA-NN-2011012pc (part and counterpart), a relatively well-preserved complete specimen (terminalia not preserved). Description: Large-sized, with body length 20.2 mm (excluding head), width of abdomen 4.9 mm . Head small with eyes projected beyond the head outline, inverted-triangular shape (length × width= 3.1 mm × 2.7 mm ), extending beyond the pronotum slightly, antennal sockets conspicuous at sides, mouthparts distinct but unclearly preserved; pronotum length 4.1 mm , width 5.2 mm , elliptical, as wide as the body; abdomen with 7 segments preserved, thick in the middle of segments, narrowing posteriorly; terminalia with forceps-like grasping cerci ( Fig. 1 B). Right forewing of holotype ( Figs. 1 , 2 ) length 14.9 mm , width 5.8 mm , conchostracan color spot present ( Fig. 1 A); left forewing length 15.4 mm , width 5.3 mm ; with small crossvein-like reticulations, wing venation rich, with 51 to 54 veins at margin ( Fig. 2 ); intercalary veins thick, distinct all over wing surface; cambered markings located near the apex of forewings, from anterior of R to the anterior of CuA, forming a crescent ( Fig. 1 A); costal area very long (about 0.4 of wing’s length); Sc thick, without branches, curved upward and longer than clavus; R slightly curved and with 22 branches (Rs undifferentiated), reaching the wing apex with many veinlets branched; M branched basally and reaching the wing apex, with 6 to 9 branches; CuA gently curved, expanded with 13 or 14 branches, with some branches ending near wing apex; CuP strongly curved and simple; clavus short, less than a third of the wing’s length; A without branches, arc bending and with 7 or 8 veins; a diagonal kink present in anal field. FIGURE 1. Arcofuzia cana Wei, Shih et Ren gen. et sp. nov. Photographs of holotype, CNU-BLA-NN-2011058. A. Habitus, Scale bar = 2 mm; B. Detail of male terminalia with forceps-like grasping cerci, Scale bar = 1 mm. Length of hind wings 13.9 –14.9 mm ; without pterostigma; with about 25 veins of remigium; Sc simple; R terminating to wing apex, differentiated into R1 with 3 branches and Rs with 10 to 12 branches; M almost straight to wing margin. Length of front femora 2.3 mm , width of front femora 0.5 mm ; length of middle femora 3.1 mm and tibiae 3.7 mm , width of middle femora 0.8 mm and tibiae 0.4 mm ; length of hind femora 4.1 mm and tibiae 4.6 mm , width of hind femora 1.0 mm and tibiae 0.4 mm ; legs gradually longer from the fore to the hind legs; hind leg with 5 spines preserved on the tibiae, length of spines 0.5–0.9 mm . Paratype ( Figs. 3 , 4 ) a relatively well-preserved complete specimen except terminalia not preserved. Large-sized, with body length 18.3 mm as preserved (excluding head), width of abdomen 4.8 mm . Head small, inverted-triangular shape (length × width= 2.6 mm × 2.3 mm ), extending beyond the pronotum slightly, antennal sockets conspicuous at sides, mouthparts distinct but unclearly preserved; pronotum length 4.5 mm , width 5.7 mm ; abdomen with 8 segments preserved, thick in the middle of segments, narrowing posteriorly. FIGURE 2. A. cana gen. et sp. nov. Line drawing of holotype, CNU-BLA-NN-2011058. Scale bar = 2 mm. Right forewing length ( Figs. 3 , 4 ) 14.9 mm , width 5.2 mm ; left forewing distorted during the fossilization process (length × width= 12.9 mm preserved × 6.0 mm preserved); with small crossvein-like reticulations ( Fig. 3 B), wing venation rich, with 39+ (left forewing) to 43 (right forewing) veins at margin; Sc thick, without branches, curved upward and longer than clavus; R slightly curved and with 16+ to 20 branches (Rs undifferentiated); M branched with 6 or 7 branches; CuA gently curved, expanded with 9 branches, with some branches ending near wing apex; A sometimes with branch ( Fig. 3 C), arc bending and with 5–6 veins. Length of front femora 1.8 mm and tibiae 1.9 mm , width of front femora 0.6 mm and tibiae 0.4 mm ; length of middle femora 2.9 mm and tibiae 3.1 mm , width of middle femora 0.9 mm and tibiae 0.4 mm ; length of hind tibiae 5.2 mm , width of hind tibiae 0.5 mm ; legs gradually longer from the fore to the hind legs; hind leg with 6 spines preserved on the tibiae, length of spines 0.4–0.9 mm . Etymology: The name cana is after the Latin word canus meaning colour of gray, referring to the colour of markings on the wings. Type locality: Daohugou Village, Wuhua Township, Ningcheng County, Inner Mongolia , China . Type horizon: Jiulongshan Formation, Middle Jurassic.