Revision of the family Acidopsidae Števčić, 2005, and the systematic position of Typhlocarcinodes Alcock, 1900, Caecopilumnus Borradaile, 1902, and Raoulia Ng, 1987, with descriptions of two new genera and five new species (Crustacea: Brachyura: Goneplacoidea) Author Ng, Peter K. L. dbsngkl@nus.edu.sg Author Rahayu, Dwi Listyo dbsngkl@nus.edu.sg text Zootaxa 2014 2014-03-03 3773 1 1 63 journal article 5890 10.11646/zootaxa.3773.1.1 6da85b06-8386-41bc-9b34-b16eba941eab 1175-5326 4909796 19F28753-B2D0-4D1F-9D47-88886F7333FD Raoulia fortis sp. nov. ( Figs. 1G, H , 5F , 6C , 20 , 21 , 41A ) Typhlocarcinodes piroculatus — Balss 1938: 73 . (not Typhlocarcinops piroculata Rathbun, 1911 ) Material examined . Holotype : male (12.0 × 9.2 mm ) ( MNHN ), station DB 8, east Aoré I., Santo, Vanuatu , 15°34.6'S 167°13.8'E , sandy patches, 12 m , coll. SANTO Expedition, 1 September 2006 . Diagnosis . Carapace width 1.3 times length (20A). Junction between frontal and supraorbital margins appears gently curved in frontal view ( Fig. 20C ). Third maxilliped with merus short, about half length of ischium; ischium quadrate, 1.1 times longer than broad ( Figs. 5F , 21B ). Ambulatory legs short; merus of last ambulatory leg 2.8 times as long as broad ( Fig. 20A ). G1 distal half slender, gently curved, with few small subdistal spinules, distal part tapering ( Fig. 21C–F ). Colour . In life, the carapace and pereopods of the type specimen are cream coloured, with patches of brown, and the setae reddish-brown ( Fig. 41A ). Etymology . The name is derived from fortis , Latin for “strong” and “stout,” alluding to the large size of the type specimen. FIGURE 19 . A–E, Raoulia piroculata ( Rathbun, 1911 ) , male (11.0 × 8.5 mm), South Africa; F–L, Raoulia galea sp. nov. , holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand. A, male abdomen; B, left third maxilliped; C, D, left G1; E, F, distal part of left G1; G, left G2. Scales: A, B = 1.0 mm; C, D, G = 0.5 mm; E, F = 0.1 mm. A–E after Barnard (1955 : fig. 16). FIGURE 20 . Raoulia fortis sp. nov. , holotype male (12.0 × 9.2 mm) (MNHN), Santo, Vanuatu. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, frontal view of carapace; D, posterior thoracic sternum; E, outer view of right chela; F, outer view of left chela. FIGURE 21 . Raoulia fortis sp. nov. , holotype male (12.0 × 9.2 mm) (MNHN), Santo, Vanuatu. A, male abdomen; B, left third maxilliped; C, D, left G1; E, F, distal part of left G1; G, left G2; H, female abdomen. Scales: A, B = 1.0 mm; C, D, G = 0.5 mm; E, F = 0.1 mm. Remarks . This new species differs from the congeners in having a relatively stouter G1 that is only gently curving and smaller spinules on the subdistal surfaces ( Fig. 21C–F ). The other species have more strongly curved G1s ( Figs. 16D–G , 19H–K , 23C–F ). The carapace and third maxilliped proportions of R. fortis sp. nov. and R. galea sp. nov. are similar, but the merus of the third maxilliped of R. fortis sp. nov. is more rounded, with the anterolateral margin more strongly convex ( Figs. 5F , 21B ) (more subovate, with the anterolateral margin only gently convex in R. galea sp. nov. , Figs. 5E , 19G ). Balss’ (1938) record of Typhlocarcinodes piroculatus ” from the Gilbert Is. (= Kiribati ) in the western Pacific was on the basis of a female specimen measuring 11.3 × 9.3 mm (no figure provided). It is probably also referable to the present new species, at least on the basis of geography. Distribution . Gilbert Is. (= Kiribati ) and Vanuatu ; 12 m .