The genus Manota Williston (Diptera: Mycetophilidae) in Melanesia and Oceania Author Hippa, Heikki text Zootaxa 2007 1502 1 44 journal article 10.5281/zenodo.177118 1513d696-cd96-4d1f-b841-2ceaae2d40f0 1175-5326 177118 Manota subspathula sp. n. ( Figs 11 A–C) A small-sized Manota . Male . Coloration . Pale brown, frons, vertex and occiput darker brown, mouthparts and legs slightly paler than thorax but apex of coxa, trochanter and base and apex of femur on legs 2 and 3 darker brown, wing unicolorous, hyaline, pale yellowish-brown, knob of haltere black. Head . Antennal flagellomere 4, Fig. 11 C. Palpomere 3 of maxillary palpus with apicomesial extension, with ca. 4 apically expanded curved sensilla; the two apical palpomeres absent on both sides of the single specimen. Thorax . Pleural setosity pattern similar to Fig. 1 D: anepisternum setose, with 24 setae; anterior basalare setose, with 6 setae; preepisternum 2 non-setose; laterotergite setose, with 26 setae; episternum 3 setose, with ca. 4 setae. Wing . Length 1.5 mm . Hypopygium ( Figs 11 A, B). Sternite 9 laterally united with gonocoxa except for the extreme posterior part, posterior margin deeply emarginated, extending to the level of the base of gonostylus, anterior margin shallowly notched, setosity similar to ventral setosity of gonocoxa. Parastylar lobe present and well exposed, with two or three setae directed obliquely posteriad, associated with weakly sclerotized lobes on its lateral side. Paraapodemal lobe not exposed in ventral view of hypopygium. The dorsal mesial margin of gonocoxa with a large lobe in the middle, bearing many blunt-ended setae at the margin and a fur-like substance on the dorsal surface, unlike other similar species. Number of juxtagonostylar setae one: it is a strong non-flattened megaseta arising from a basal body, which is slightly shorter than half the length of the megaseta. Gonostylus elongate, widening towards the slightly bilobed apex, with two long apical setae, with a row of dorsomesial setae diverging from the other rather weak setosity. Tegmen unusually shaped, with very strong lateral shoulders. Hypoproct long, extending posteriorly nearly as far as the apex of gonostylus, with ca. 20 scattered ventral setae. Cerci separate. Female and preimaginal stages unknown. FIGURE 11 . Manota subspathula sp. n. (holotype): A . Hypopygium, dorsal view. B. Hypopygium, ventral view. C. Antennal flagellomere 4, lateral view. Scale for A and B 0.10 mm, for C 0.05 mm. Discussion. M. subspathula is similar to M. spathula but can be distinguished e.g. by the following characters: 1) the posterior margin of sternite 9 is notched, not convex, 2) the large lobe at the middle of the dorsal mesial margin of gonocoxa has a fur- or velvet-like substance on it dorsal surface, bare in M. spathula , 3) the juxtagonostylar megaseta is narrow, greatly expanded in M. spathula , 4) the long apical setae of gonostylus are wide apart, with one at the apicolateral corner and the other at the apicomesial corner, in M. subspathula both the setae are close to each other at the apicolateral corner, and 5) the lateral shoulders of tegmen are much more strongly pronounced. Both species are similar to M. hirsuta and to many species outside the Melanesian and Oceanian region. For further discussion, see under M. hirsuta . Types . Holotype . Male , Papua New Guinea , Madang province, Baitabag village, Kau Wildlife Area, 50 m a. s. l., S 5o 08’ E 145o 46’ , primary forest, Malaise trap 8– 21.11.1999 , Lukáš Č iżek leg. (in NRM).