Review of Mycetobia Meigen, 1818 (Diptera, Anisopodidae) in the Eocene ambers Author Wojtoń, Maciej Author Kania, Iwona Author Krzemiński, Wiesław text Zootaxa 2019 2019-01-10 4544 1 1 40 journal article 27687 10.11646/zootaxa.4544.1.1 68749531-5011-4ec9-93e5-f48416d0843f 1175-5326 2618075 BE4670E8-0A88-4838-84BC-F4AB6E2F797B Mycetobia hansi sp. nov. ( Figs 14–15 ) urn:lsid:zoobank.org:act: B8B4B962-E5C4-4989-8256-2C6F348154C0 Diagnosis. Flagellomeres 1–7 wider than long, flagellomeres 8–13 longer than wide, became more elongated to the apex of antenna, up to 2x as long as wide; last flagellomere at most 2x as long as wide; last palpomere elongated 3x as long as wide, tapered at apex, longer than second one; Sc ending before Rs bifurcation level; R 1 apex before M 1+2 bifurcation level; the distance between R 1 and R 2+3 apices approximately 4x as long as the distance between Sc and R 1 apices; R 2+3 2.5x as long as Rs; the distance between Sc and R 1 apices approximately 4x as long as the distance between R 1 and R 2+3 apices; M 1+2 shorter than M 1 ; fork of M 1+2 ; the distance between M 1 and M 2 apices slightly wider than the distance between R 4+5 and M 1 apices; the distance between M 2 and M 3+4 apices longer than the distance between M 1 and M 2 ; R 2+3 ending opposite bifurcation of M 1+2 level, before half the length R 4+5 level; only prescutellar and apical scutellar setae thick, but not very elongated. Etymology. The species name is dedicated to Hans Werner Hoffeins from Hamburg , Germany , the owner of amber inclusions collection and the expert in Baltic amber inclusions. Type material. Holotype : No. 1258-4 (female), Baltic amber, (coll. Ch. & H. W. Hoffeins). Description. Body ( Fig. 15A ) 3.28 mm long, pale brown with dark brown thorax. Head ( Figs 15A, B ): 0.62 mm high; ocelli present and form equilateral triangle. Antenna 1.06 mm long ( Figs 14A ; 15A, B ), 14 flagellomeres, more or less elongated, scape and pedicel comparatively short and wide, wider than long, with a few very elongated setae in distal part, sometimes longer than segments bearing them; last flagellomere tapered to the end, longer than penultimate one, approximately twice as long as wide; flagellomeres covered by comparatively elongated sparse and thick setae, but setae much shorter than segments bearing them; additionally very short setae on all flagellomeres, last flagellomere with a few not very elongated apical setae; palpus maxillaris 4-segmented, slender, first palpomere 0.04 mm long cylindrical, second palpomere not very elongated, 0.09 mm long, only slightly longer than wide, widened in the midlenght, third small, short, 0.03 mm long, approximately as long as wide, last one elongated, 0.10 mm long, tapered at apex ( Figs 14B ; 15B ) elongated and short setae on all palpomeres. FIGURE 14. Mycetobia hansi sp. nov. holotype, No. 1258-4 (female), (coll. Ch & H. W. Hoffeins). A. antenna; B. palpus maxillaris; C. wing venation; D. diagram of relation between the length of legs: fore leg (F), middle leg (M), hind leg (H); point indicate position of tibial spurs. Abbreviations: SC—scape; PED—pedicel; f1–f14—flagellomeres; I–IV numbers of tarsomeres. FIGURE 15. Mycetobia hansi sp. nov. holotype, No. 1258-4 (female), (coll. Ch & H. W. Hoffeins). A. habitus, latero-dorsal view; B. enlarged view of head with antenna and palpus maxillaris, latero-ventral view; C. wing. Thorax: prescutellar setae and apical scutellar setae very elongated; acrostichal setulae not very elongated; wing ( Figs 14C ; 15C ): 3.08 mm long, 1.03 mm wide; Sc comparatively short, slightly arched, ending at 1/2 of Rs level, Rs 0.33 mm long; Rs not very short, R 2+3 0.66 mm long; R 1 ending before midlength, at R 4+5 level; M 1+2 approximately as long as M 2 and shorter than M 1 , M 1+2 0.74 mm long, A 1 well developed, connected with wing edge; A 2 short and slender. Halter not very elongated with elongated stem and widened knob. Legs ( Figs 15A ; 14D ): fore, middle and hind tibia with two unequal tibial spurs; tarsus comparatively elongated with last segment only slightly longer than penultimate one and not very elongated, rather slender claw and almost invisible empodium. Apical comb well visible on fore and hind legs. Tarsus of fore leg shorter than 0.3 the length of leg. Fore coxa 0.59 mm , trochanter 0.08 mm , femur 0.89 mm , tibia 0.80 mm , tarsus 1.96 mm long (0.38/0.12/0.11/0.01/ 0.02). Middle coxa 0.42 mm long, trochanter 0.12 mm long, femur 0.57 mm long, tibia 0.79 mm , tarsus 0.95 mm (0.55/0.15/0.09/0.09/0.09). The length of hind coxa 0.42 mm , trochanter 0.12 mm , femur 1.02 mm , tibia 0.88 mm , tarsus 1.22 mm (0.67/0.18/0.13/0.11/0.13). Abdomen: female terminalia not very elongated, cerci not very large Comparison. In contrast to other species known from the Baltic amber and described herein vein M 1+ 2 in M. hansi sp. nov. is distinctly shorter than M 1 . There are also some other differences in wing venation; in contrast to M. silvia sp. nov. , where Sc ending beyond Rs bifurcation level, R 1 apex is positioned beyond M 1+2 bifurcation level, in M. hansi sp. nov. Sc ending before Rs bifurcation level and R 1 apex is positioned before M 1+2 bifurcation level. Moreover, in contrast to M. christelae sp. nov. , where R 2+3 is 3x as long as Rs, R 2+3 ending beyond half the length R 4+5 level, in M. hansi sp. nov. R 2+3 is 2.5x as long as Rs, R 2+3 ending before half the length R 4+5 level. More differences of wing venation is given in Table 2. M . hansi sp. nov. differ distinctly from M. silvia sp. nov. by morphology of antenna. In M. hansi sp. nov. flagellomeres 1–7 are wider than long, flagellomeres 8–13 are longer than wide, became more elongated to the apex of antenna, the last one is at most 2x as long as wide, while in M. silvia sp. nov. flagellomeres are cylindrical, becoming narrowed and more elongated to the apex of antenna, only flagellomeres 1–3 are as long as wide, flagellomeres 4–7 are more than 1x to 1.5x as long as wide, flagellomeres 8–13 are elongated 2x or more longer than wide, the last one is 3x as long as wide. Prescutellar and apical scutellar setae are thick, but not very elongated in M. hansi sp. nov. , not like in M. szwedoi sp. nov. where are very elongated and thick.