Testing the phylogenetic utility of morphological character systems, with a revision of Creophilus Leach (Coleoptera: Staphylinidae) Author Clarke, Dave J. text Zoological Journal of the Linnean Society 2011 Zool. J. Linn. Soc. 2011-10-25 163 3 723 812 http://dx.doi.org/10.1111/j.1096-3642.2011.00725.x journal article 10.1111/j.1096-3642.2011.00725.x 0024-4082 5441985 FBFE9195-BE04-4AFE-9417-6E38BCE6AB84 THE CREOPHILUS COMPLEX Adult diagnosis: Species of the Creophilus complex can be recognized by the combination of: disc of head and pronotum glabrous; ventroscapal macrosetae absent or reduced; superior marginal pronotal carina separate from inferior marginal carina along entire length, effaced apically ( Fig. 2E , il , sl , Fig. 9J , right and middle arrows); inferior marginal pronotal carina continuous with anterior marginal carina ( Fig. 9J , left and middle arrows); mesepimeron with single marginal ridge ( Fig. 9K , arrow); male sternite IX subtruncate to shallowly emarginate apically, produced into elongate process basally ( Fig. 3Q ); median lobe of aedeagus with large apicolateral sclerites ( Fig. 3 , as ); internal sac without ventral basomedian sclerotized strip (as in Fig. 3K , bms ). Remarks: The following taxa are here included in the Creophilus complex: Creophilus (12 species), Hadrotes crassus , Ht. wakefieldi , Hadropinus fosso r, Liusus (two species), Thinopinus pictus .