<document id="EE6DCFA263C0D214CAA012C97F2FFC8F" ID-DOI="10.5281/zenodo.3678492" ID-GBIF-Dataset="8e18452b-1d9d-462e-bac8-8c86eeb4ddc3" ID-ISSN="1942-1354" ID-Zenodo-Dep="3678492" ID-ZooBank="930EAB11-37FA-41B0-980A-1A4736527842" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1582295086673" checkinUser="felipe" docAuthor="Hielkema, Auke J. &amp; Hielkema, Meindert A." docDate="2019" docId="03D7FF1CFFC2FFE4FF48F950302BFCDB" docLanguage="en" docName="InsectaMundi.0732.1-306.pdf.imf" docOrigin="Insecta Mundi 732 (732)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Athyreus biarmatus Howden and Martinez 1978" docType="treatment" docVersion="12" lastPageNumber="8" masterDocId="FFEE8764FFCBFFEDFF88FB6E317CFFF0" masterDocTitle="An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas" masterLastPageNumber="306" masterPageNumber="1" pageNumber="8" updateTime="1698721431862" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="487B60722620D8E9BE15A197596F92FF">An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas</mods:title>
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<paragraph id="8BC14E0AFFC2FFE4FF48F9503215FDA6" blockId="9.[192,873,573,598]" box="[192,873,573,598]" pageId="9" pageNumber="8">
<heading id="D089F966FFC2FFE4FF48F9503215FDA6" bold="true" box="[192,873,573,598]" fontSize="10" level="2" pageId="9" pageNumber="8" reason="3">
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§ 
<taxonomicName id="4C7E3589FFC2FFE4FF5DF9503215FDA6" authority="Howden and Martinez, 1978" authorityName="Howden and Martinez" authorityYear="1978" box="[213,873,573,598]" class="Insecta" family="Bolboceratidae" genus="Athyreus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="biarmatus">
<emphasis id="B90A9218FFC2FFE4FF5DF950309FFDA6" bold="true" box="[213,483,574,598]" italics="true" pageId="9" pageNumber="8">Athyreus biarmatus</emphasis>
<bibRefCitation id="EFEF33FBFFC2FFE4FE63F9503215FDA6" author="Howden and Martinez" box="[491,873,573,598]" firstAuthor="Howden" pageId="9" pageNumber="8" pagination="8 - 19" refId="ref214741" refString="Martinez, A. 1978. Algunos Cyclocephalini neotropicales nuevos (Col. Scarab. Dynastinae). Acta Cientifica, Serie Entomologia 12: 8 - 19." type="journal article" year="1978">Howden and Martínez, 1978</bibRefCitation>
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</emphasis>
</heading>
</paragraph>
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<paragraph id="8BC14E0AFFC2FFE4FF60F90A3559FD8B" blockId="9.[232,1061,612,635]" box="[232,1061,612,635]" pageId="9" pageNumber="8">
<treatmentCitationGroup id="AB6E6924FFC2FFE4FF60F90A3559FD8B" box="[232,1061,612,635]" pageId="9" pageNumber="8">
<treatmentCitation id="0ADF681BFFC2FFE4FF60F90A3259FD8B" author="Howden and Martinez" box="[232,805,612,635]" page="35" pageId="9" pageNumber="8" year="1978">
<taxonomicName id="4C7E3589FFC2FFE4FF60F90A3259FD8B" authority="Howden and Martinez 1978: 35" authorityName="Howden and Martinez" authorityPageNumber="35" authorityYear="1978" box="[232,805,612,635]" class="Insecta" family="Bolboceratidae" genus="Athyreus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="biarmatus">
<emphasis id="B90A9218FFC2FFE4FF60F90A30B8FD89" box="[232,452,612,633]" italics="true" pageId="9" pageNumber="8">Athyreus biarmatus</emphasis>
<bibRefCitation id="EFEF33FBFFC2FFE4FE42F90B3259FD8B" author="Howden and Martinez" box="[458,805,613,635]" firstAuthor="Howden" pageId="9" pageNumber="8" pagination="8 - 19" refId="ref214741" refString="Martinez, A. 1978. Algunos Cyclocephalini neotropicales nuevos (Col. Scarab. Dynastinae). Acta Cientifica, Serie Entomologia 12: 8 - 19." type="journal article" year="1978">Howden and Martínez 1978: 35</bibRefCitation>
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(
<collectingCountry id="F3690E9AFFC2FFE4FCBCF90B3562FD8B" box="[820,1054,613,635]" name="Trinidad and Tobago" pageId="9" pageNumber="8">Trinidad and Tobago</collectingCountry>
)
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</paragraph>
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Distribution. 
<collectingCountry id="F3690E9AFFC2FFE4FEF6F9E53370FD53" box="[382,524,651,675]" name="Venezuela" pageId="9" pageNumber="8">Venezuela</collectingCountry>
(EO):
</emphasis>
<bibRefCitation id="EFEF33FBFFC2FFE4FDE2F9E53228FD53" author="Howden" box="[618,852,651,675]" firstAuthor="Howden" journalOrPublisher="The Coleopterists Bulletin" pageId="9" pageNumber="8" pagination="339 - 354" part="53" refId="ref208514" refString="Howden, H. F. 1999. New species of Central and South American Athyreini (Coleoptera: Scarabaeidae: Geotrupinae). The Coleopterists Bulletin 53 (4): 339 - 354." title="New species of Central and South American Athyreini (Coleoptera: Scarabaeidae: Geotrupinae)" type="journal article" year="1999">Howden 1999: 345</bibRefCitation>
(EO). 
<emphasis id="B90A9218FFC2FFE4FC24F9E53578FD53" bold="true" box="[940,1028,651,675]" pageId="9" pageNumber="8">Other:</emphasis>
<bibRefCitation id="EFEF33FBFFC2FFE4FB87F9E534DCFD53" author="Howden and Martinez" box="[1039,1440,651,675]" firstAuthor="Howden" pageId="9" pageNumber="8" pagination="8 - 19" refId="ref214741" refString="Martinez, A. 1978. Algunos Cyclocephalini neotropicales nuevos (Col. Scarab. Dynastinae). Acta Cientifica, Serie Entomologia 12: 8 - 19." type="journal article" year="1978">Howden and Martínez 1978: 35</bibRefCitation>
(Trinidad and Tobago); 
<bibRefCitation id="EFEF33FBFFC2FFE4FE69F9C533BEFD33" author="Howden" box="[481,706,683,707]" firstAuthor="Howden" journalOrPublisher="The Coleopterists Bulletin" pageId="9" pageNumber="8" pagination="339 - 354" part="53" refId="ref208514" refString="Howden, H. F. 1999. New species of Central and South American Athyreini (Coleoptera: Scarabaeidae: Geotrupinae). The Coleopterists Bulletin 53 (4): 339 - 354." title="New species of Central and South American Athyreini (Coleoptera: Scarabaeidae: Geotrupinae)" type="journal article" year="1999">Howden 1999: 345</bibRefCitation>
(Trinidad and Tobago).
</paragraph>
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<emphasis id="B90A9218FFC2FFE4FF48F9BD3074FD1B" bold="true" box="[192,264,723,747]" pageId="9" pageNumber="8">Note.</emphasis>
<bibRefCitation id="EFEF33FBFFC2FFE4FE98F9BD337FFD1B" author="Howden" box="[272,515,723,747]" firstAuthor="Howden" journalOrPublisher="The Coleopterists Bulletin" pageId="9" pageNumber="8" pagination="339 - 354" part="53" refId="ref208514" refString="Howden, H. F. 1999. New species of Central and South American Athyreini (Coleoptera: Scarabaeidae: Geotrupinae). The Coleopterists Bulletin 53 (4): 339 - 354." title="New species of Central and South American Athyreini (Coleoptera: Scarabaeidae: Geotrupinae)" type="journal article" year="1999">Howden (1999: 345)</bibRefCitation>
suspects 
<taxonomicName id="4C7E3589FFC2FFE4FDF2F9BD3211FD1B" authorityName="Howden and Martinez" authorityYear="1978" box="[634,877,723,747]" class="Insecta" family="Bolboceratidae" genus="Athyreus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="biarmatus">
<emphasis id="B90A9218FFC2FFE4FDF2F9BD3211FD1B" box="[634,877,723,747]" italics="true" pageId="9" pageNumber="8">Athyreus biarmatus</emphasis>
</taxonomicName>
to be dimorphic in males, with the other form being 
<taxonomicName id="4C7E3589FFC2FFE4FE80F99D3325FCFB" authority="Westwood, 1848" authorityName="Westwood" authorityYear="1848" box="[264,601,755,779]" class="Insecta" family="Bolboceratidae" genus="Athyreus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B90A9218FFC2FFE4FE80F99D30F3FCFB" box="[264,399,755,779]" italics="true" pageId="9" pageNumber="8">A. armatus</emphasis>
<bibRefCitation id="EFEF33FBFFC2FFE4FE1CF99D3325FCFB" author="Westwood" box="[404,601,755,779]" firstAuthor="Westwood" pageId="9" pageNumber="8" pagination="386 - 387" refId="ref229867" refString="Westwood, J. O. 1848. Descriptions of some new species of Athyreus MacL., a genus of lamellicorn beetles. Annals and Magazine of Natural History, including Zoology, Botany, and Geology 2 (1) (5): 386 - 387." type="journal article" year="1848">Westwood, 1848</bibRefCitation>
</taxonomicName>
(p. 386). In case they belong to the same species, the name 
<taxonomicName id="4C7E3589FFC2FFE4FA91F99D34DCFCFB" authorityName="Westwood" authorityYear="1848" box="[1305,1440,755,779]" class="Insecta" family="Bolboceratidae" genus="Athyreus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B90A9218FFC2FFE4FA91F99D34DCFCFB" box="[1305,1440,755,779]" italics="true" pageId="9" pageNumber="8">A. armatus</emphasis>
</taxonomicName>
has priority.
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