Order Primates Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1 111 184 book chapter 0-8018-8221-4 10.5281/zenodo.7316519 Saguinus Hoffmannsegg 1807 Saguinus Hoffmannsegg 1807 , Mag. Ges. Naturf. Fr., 1: 101 . Type Species: Saguinus ursula Hoffmannsegg 1807 Synonyms: Hapanella Gray 1870 ; Leontocebus Wagner 1840 ; Marikina Lesson 1840 ; Midas E. Geoffroy St. Hilaire 1812 ; Mystax Gray 1870 ; Oedipomidus Reichenbach 1862 ; Oedipus Lesson 1840 ; Seniocebus Gray 1870 ; Tamarin Gray 1870 ; Tamarinus Trouessart 1904 . Species and subspecies: 17 species with 21 subspecies: Species Saguinus bicolor Spix 1823 Species Saguinus fuscicollis Spix 1823 Subspecies Saguinus fuscicollis subsp. fuscicollis Spix 1823 Subspecies Saguinus fuscicollis subsp. avilapiresi Hershkovitz 1966 Subspecies Saguinus fuscicollis subsp. cruzlimai Hershkovitz 1966 Subspecies Saguinus fuscicollis subsp. fuscus Lesson 1840 Subspecies Saguinus fuscicollis subsp. illigeri Pucheran 1845 Subspecies Saguinus fuscicollis subsp. leucogenys Gray 1866 Subspecies Saguinus fuscicollis subsp. lagonotus Jiménez de la Espada 1870 Subspecies Saguinus fuscicollis subsp. nigrifrons I. Geoffroy 1851 Subspecies Saguinus fuscicollis subsp. primitivus Hershkovitz 1977 Subspecies Saguinus fuscicollis subsp. weddelli Deville 1849 Species Saguinus geoffroyi Pucheran 1845 Species Saguinus graellsi Jimenez de la Espada 1870 Species Saguinus imperator Goeldi 1907 Subspecies Saguinus imperator subsp. imperator Goeldi 1907 Subspecies Saguinus imperator subsp. subgrisescens Lönnberg 1940 Species Saguinus inustus Schwartz 1951 Species Saguinus labiatus E. Geoffroy 1812 Subspecies Saguinus labiatus subsp. labiatus E. Geoffroy 1812 Subspecies Saguinus labiatus subsp. rufiventer Gray 1843 Subspecies Saguinus labiatus subsp. thomasi Goeldi 1907 Species Saguinus leucopus Günther 1876 Species Saguinus martinsi Thomas 1912 Subspecies Saguinus martinsi subsp. martinsi Thomas 1912 Subspecies Saguinus martinsi subsp. ochraceus Hershkovitz 1966 Species Saguinus melanoleucus Miranda Ribeiro 1912 Species Saguinus midas ( Linnaeus 1758 ) Species Saguinus mystax Spix 1823 Subspecies Saguinus mystax subsp. mystax Spix 1823 Subspecies Saguinus mystax subsp. pluto Lönnberg 1926 Species Saguinus niger (E. Geoffroy 1803 ) Species Saguinus nigricollis Spix 1823 Subspecies Saguinus nigricollis subsp. nigricollis Spix 1823 Subspecies Saguinus nigricollis subsp. hernandezi Hershkovitz 1982 Species Saguinus oedipus ( Linnaeus 1758 ) Species Saguinus pileatus I. Geoffroy and Deville 1848 Species Saguinus tripartitus Milne-Edwards 1878 Discussion: See Hershkovitz (1977:601-603) . For a summary of interrelationships between species, using external, cranial, vocalization and DNA evidence, see Groves (2001 c :137-8) , who divided the genus into the following species groups: (1) S. midas group ( midas , niger ), (2) S. nigricollis group ( nigricollis , graellsi , fuscicollis , melanoleucus , tropartitus ), (3) S. mystax group ( mystax , pileatus , labiatus , imperator ), (4) S. bicolor group ( bicolor , martinsi ), (5) S. oedipus group ( oedipus , geoffroyi , leucopus ), (6) S. inustus group ( inustus only).