Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar Author Kavanaugh, David H. Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; & Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org) Author Rainio, Johanna Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com text Proceedings of the California Academy of Sciences 2016 2016-04-29 63 7 201 268 journal article 10.5281/zenodo.13799439 0068-547X 13799439 Thysanotus bimaculatoides Kavanaugh and Rainio , sp. nov. Figures 17–18 TYPE MATERIAL .— Holotype ( Figs.16A–B ), a male, in CAS, labeled: “ MADAGASCAR Ranomafana NP. Mangevo plot 1600m 20.4.2005 ”/ “240” [handwritten label]/ “ HOLOTYPE FIGURE 17. A-D. Digital images of holotype male of Thysanotus bimaculatoides Kavanaugh & Rainio sp. nov. A–D. A Habitus, Dorsal aspect; B. Labels. Scale line = 1.0 mm. FIGURE 18. Digital images of aedeagus of male genitalia. A, C. Left lateral aspect; B, D. Dorsal aspect. A–B. Holotype male of Thysanotus bimaculatoides Kavanaugh & Rainio sp. nov. ; B, D. Thysanotus spinosus Alluaud (from Ranomafana National Park). Scale lines = 0.5 mm. Thysanotus bimaculatoides Kavanaugh & Rainio sp. n. 2015” [red label]. Paratypes (total of 2): 1 female (in MNHN ) labeled “ MADAGASCAR Ranomafana NP. Mangevo plot 1600m 22.4.2005 ”/ “240” [handwritten label]; 1 female (in CAS ) labeled “CASENT 8069538”/ “ MADAGASCAR : Province Fianarantsoa , Forêt d’Ambalagoavy Nord , Ikonga , Abatombe Nov-2000 ”/ “ 21°49’39”S , 47°20’20”E Calif. Acad. of Sciences colls: R. Harin’Hala & M.E. Irwin , malaise trap 625m MA-01-12-01”. Both paratypes also bear the following label: “ PARATYPE Thysanotus bimaculatoides Kavanaugh & Rainio sp. n. 2015” [yellow label]. TYPE LOCALITY .— Madagascar , Fianarantsoa Province , Ranomafana National Park . DERIVATION OF SPECIES NAME .— The species epithet, bimaculatoides , is an adjective derived from the Latin words, bis , meaning two, and macula , meaning spot or spotted, and the Greek word eidos , meaning like. The name refers to the similarity between members of this species and those of Thysanotus bimaculatus Deuve (2010) . RECOGNITION .— Size slightly less than average for genus, SBL of male = 5.7 mm , of females = 5.3 mm . Members of this species ( Fig. 17A ) key to Thysanotus bimaculatus Deuve (2010) in Deuve’s key based on the following combination of features: elytra shiny, intervals smooth, elytral microsculpture indistinct (comprised of faintly impressed microlines and markedly transverse sculpticells, each elytron with a large, pale macula near the base on intervals 2 to 4, also extended partially onto interval 1 and to scutellum at basomedial part of macula, elytral apex bluntly round- ed, not toothed. Members of T. bimaculatoides differ from the holotype of T. bimaculatus in having slightly larger body size (SBL of holotype female of T. bimaculatus = 4.9 mm ), pronotum slightly narrower, ratio of PW/PL = 1.1 (ratio of PW/PL = 1.2 in T. bimaculatus ), narrower basally with subbasal sinuation of the lateral margin slightly longer and deeper and hind angles sharp- er, rectangular to distinctly acute (compare with Deuve 2010 , Fig. 17 ). Deuve did not have a male specimen of T. bimaculatus , so we cannot compare male genitalia of these two species. However, for future comparative purposes, we provide digital photographs of the genitalia ( Figs. 18A,C ) of the holotype of T. bimaculatoides as well as those of a male of Thysanotus spinosus ( Alluaud 1936c ) ( Figs. 18B,D ) [males of this species previously unknown], to complement the illustrations of other Thysanotus species provided by Jeannel (1949) and Deuve (2010) . GEOGRAPHICAL DISTRIBUTION .— At present, known only from two localities, 52 km apart, in central Fianarantsoa Province . Both localities are on the eastern slope off the central plateau. HABITAT DISTRIBUTION .— The holotype and one paratype were collected by hand in the Mangevo area , about 1600 m distance in from the edge of montane rainforest, at an elevation of 1600 m . The other paratype was collected in a malaise trap in montane rainforest at an elevation of 650 m .