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<mods:title id="FBF9DAD3AC04DFF84CD941AF2B88C5B5">An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas</mods:title>
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<emphasis id="B90A9218FFC2FFE4FF48FD843017F8F2" bold="true" box="[192,363,1770,1794]" italics="true" pageId="9" pageNumber="8">Neoathyreus</emphasis>
(
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) 
<emphasis id="B90A9218FFC2FFE4FDC0FD85339CF8F3" bold="true" box="[584,736,1771,1795]" italics="true" pageId="9" pageNumber="8">guyanensis</emphasis>
<bibRefCitation id="EFEF33FBFFC2FFE4FD6FFD8532D9F8F3" author="Howden" box="[743,933,1771,1795]" firstAuthor="Howden" journalOrPublisher="Zootaxa" pageId="9" pageNumber="8" pagination="37 - 52" part="1137" refId="ref208622" refString="Howden, H. F. 2006. New species and a new subgenus of South American Neoathyreus Howden and Martinez (Coleoptera: Geotrupidae: Athyreini). Zootaxa 1137: 37 - 52." title="New species and a new subgenus of South American Neoathyreus Howden and Martinez (Coleoptera: Geotrupidae: Athyreini)" type="journal article" year="2006">Howden, 2006</bibRefCitation>
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<emphasis id="B90A9218FFC2FFE4FF60FC7F300DF8D6" box="[232,369,1809,1830]" italics="true" pageId="9" pageNumber="8">Neoathyreus</emphasis>
(
<emphasis id="B90A9218FFC2FFE4FE08FC7F336AF8D6" box="[384,534,1809,1830]" italics="true" pageId="9" pageNumber="8">Mesoathyreus</emphasis>
) 
<emphasis id="B90A9218FFC2FFE4FDADFC7C33DCF8D7" box="[549,672,1810,1831]" italics="true" pageId="9" pageNumber="8">guyanensis</emphasis>
<bibRefCitation id="EFEF33FBFFC2FFE4FD2FFC7F321AF8D7" author="Howden" box="[679,870,1809,1831]" firstAuthor="Howden" journalOrPublisher="Zootaxa" pageId="9" pageNumber="8" pagination="37 - 52" part="1137" refId="ref208622" refString="Howden, H. F. 2006. New species and a new subgenus of South American Neoathyreus Howden and Martinez (Coleoptera: Geotrupidae: Athyreini). Zootaxa 1137: 37 - 52." title="New species and a new subgenus of South American Neoathyreus Howden and Martinez (Coleoptera: Geotrupidae: Athyreini)" type="journal article" year="2006">Howden 2006: 50</bibRefCitation>
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(
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Distribution. 
<collectingCountry id="F3690E9AFFC2FFE4FEF2FC563330F8BF" box="[378,588,1847,1872]" name="French Guiana" pageId="9" pageNumber="8">French Guiana</collectingCountry>
:
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; 
<bibRefCitation id="EFEF33FBFFC2FFE4FCB2FC56357EF8A0" author="Boilly" box="[826,1026,1848,1872]" firstAuthor="Boilly" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="9" pageNumber="8" pagination="19 - 31" part="4" refId="ref194293" refString="Boilly, O. 2011 b. Les Bolboceratidae de Guyane (Coleoptera, Scarabaeoidea). Contribution a l'etude des coleopteres de Guyane 4: 19 - 31. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Les Bolboceratidae de Guyane (Coleoptera, Scarabaeoidea)" type="journal article" year="2011" yearSuffix="b">Boilly 2011b: 24</bibRefCitation>
.
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Subgenus 
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<emphasis id="B90A9218FFC1FFE7FEB7FBB7335DFF01" bold="true" box="[319,545,217,241]" italics="true" pageId="10" pageNumber="9">NEOATHYREUS</emphasis>
<bibRefCitation id="EFEF33FBFFC1FFE7FDA1FBB732DBFF01" author="Howden and Martinez" box="[553,935,217,241]" firstAuthor="Howden" journalOrPublisher="The Canadian Entomologist" pageId="10" pageNumber="9" pagination="345 - 352" part="95" refId="ref208846" refString="Howden, H. F., and A. Martinez. 1963. The new tribe Athyreini and its included genera (Coleoptera: Scarabaeidae, Geotrupinae). The Canadian Entomologist 95 (4): 345 - 352." title="The new tribe Athyreini and its included genera (Coleoptera: Scarabaeidae, Geotrupinae)" type="journal article" year="1963">Howden and Martínez, 1963</bibRefCitation>
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<taxonomicName id="4C7E3589FFC1FFE7FF60FA6E339EFEE6" authority="Howden and Martinez 1963: 350" authorityName="Howden and Martinez" authorityPageNumber="350" authorityYear="1963" box="[232,738,256,278]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B90A9218FFC1FFE7FF60FA6E300CFEE5" box="[232,368,256,277]" italics="true" pageId="10" pageNumber="9">Neoathyreus</emphasis>
<bibRefCitation id="EFEF33FBFFC1FFE7FEF0FA6E339EFEE6" author="Howden and Martinez" box="[376,738,256,278]" firstAuthor="Howden" journalOrPublisher="The Canadian Entomologist" pageId="10" pageNumber="9" pagination="345 - 352" part="95" refId="ref208846" refString="Howden, H. F., and A. Martinez. 1963. The new tribe Athyreini and its included genera (Coleoptera: Scarabaeidae, Geotrupinae). The Canadian Entomologist 95 (4): 345 - 352." title="The new tribe Athyreini and its included genera (Coleoptera: Scarabaeidae, Geotrupinae)" type="journal article" year="1963">Howden and Martínez 1963: 350</bibRefCitation>
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