Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain) Author Oliver, Joan Daniel Alcorisa 83 - 12 C, 28043 Madrid, Spain. joandanieloliver @ hotmail. com Author Gofas, Serge Departamento de Biología Animal, Universidad de Málaga, E- 29071 Málaga, Spain. Author Urgorri, Victoriano Estación de Bioloxía Mariña da Graña, REBUSC, Universidade de Santiago de Compostela, Spain. Author Díaz-Agras, Guillermo Estación de Bioloxía Mariña da Graña, REBUSC, Universidade de Santiago de Compostela, Spain. Author Templado, José Museo Nacional de Ciencias Naturales (CSIC), José Gutiérrez Abascal 2, 28006 Madrid, Spain. text Zootaxa 2022 2022-10-18 5196 1 1 45 journal article 170600 10.11646/zootaxa.5196.1.1 e1042177-7e89-413a-b10b-3803d000d20d 1175-5326 7224080 1B12892E-D68F-4BFE-8E9A-7F19E220E73D Alvania testae (Aradas & Maggiore, 1844) ( Figs. 5 e–g ) Material examined. ( nearly 2000 shells, most of them with soft parts, in 33 samples): SPAIN • 1 s; 43° 40.165′N , 008° 43.697′W to 43° 40.513′N, 008° 43.159′W; 204–209 m ; 08–15 SeP. 2002 ; DIVA-Artabria I DRN-200 • >80 sPc; 43° 35.451′N , 008° 34.432′W to 43° 34.810′N, 008° 35.407′W; 153– 151 m ; 08–15 SeP. 2002 ; DIVA-Artabria I EBS-150 • 40 spc; 43° 40.192′N , 008° 43.760′W to 43° 40.943′N, 008° 42.366′W; 207–212 m ; 08–15 SeP. 2002 ; DIVA-Artabria I EBS-200 • 10 sPc; 43° 41.689′N , 008° 45.195′W to 43° 42.556′N, 008° 44.226′W; 298–303 m ; 08–15 SeP. 2002 ; DIVA-Artabria I EBS-300 • 1 spc (juv); 43° 53.575′N , 008° 56.868′W to 43° 54.015′N, 008° 56.959′W; 965–974 m ; 10–20 SeP. 2003 ; DIVA-Artabria I DRN-1000 • > 80 sPc; 43° 34.116′N , 008° 36.535′W to 43° 34.689′N, 008° 35.589′W; 149– 148 m ; 10–20 Sep. 2003 ; DIVA-Artabria I AT-150 • > 200 spc; 43° 34.127′N , 008° 36.562′W to 43° 34.820′N, 008° 35.585′W; 152– 149 m ; 10–20 Sep. 2003 ; DIVA-Artabria I EBS-150 • > 50 spc; 43° 40.250′N , 008° 43.755′W to 43° 40.760′N, 008° 42.120′W; 207– 197 m ; 10–20 Sep. 2003 ; DIVA-Artabria I EBS-200 • > 300 spc; 43° 41.590′N , 008° 45.328′W to 43° 42.396′N, 008° 44.286′W; 301– 301 m ; 10–20 SeP. 2003 ; DIVA-Artabria I EBS-300 • 1 sh; 43° 42.348′N , 008° 45.889′W to 43° 43.269′N, 008° 45.289′W; 344–354 m ; 10–20 SeP. 2003 ; DIVA-Artabria I EBS-350 • 30 spc + 200 sh; 43° 29.412′N , 008° 28.362′W to 43° 29.779′N, 008° 28.240′W; 110–111 m ; 17–28 SeP. 2004 ; VERTIDOS GA-AT-110 • 50 sPc; 43° 32.026′N , 008° 37.520′W to 43° 32.806′N, 008° 35.993′W; 150–151 m ; 17–28 SeP. 2004 ; VERTIDOS GA-AT-150 • 1 spc (juv); 43° 38.105′N , 009° 02.006′W to 43° 37.738′N, 009° 00.905′W; 836– 826 m ; 17–28 SeP. 2004 ; VERTIDOS GA-AT-800 • 30 sPc; 42° 29.455′N , 009° 17.472′W to 42° 31.292′N, 009° 19.660′W; 149– 147 m ; 17–28 SeP. 2004 ; VERTIDOS AG-AT-150 • >40 sPc; 42° 47.353′N , 009° 25.161′W to 42° 44.866′N, 009° 25.913′W; 147– 146 m ; 17–28 SeP. 2004 ; VERTIDOS CA-AT-150 • 30 spc; 43° 31.970′N , 008° 37.670′W to 43° 32.697′N, 008° 3.171′W; 151– 151 m ; 17–28 SeP. 2004 ; VERTIDOS GA-EBS-150 • 25 spc; 43° 36.222′N , 008° 52.845′W to 43° 36.797′N, 008° 52.112′W; 201– 201 m ; 17–28 Sep. 2004 ; VERTIDOS GA-EBS-200 • >800 spc; 42° 30.391′N , 009° 19.517′W to 42° 29.428′N, 009° 17.924′W; 147–148 m ; 17–28 Sep. 2004 ; VERTIDOS AG-EBS-150 • >50 spc; 42° 31.176′N , 009° 23.380′W to 42° 32.132′N, 009° 24.151′W; 242–248 m ; 17–28 Sep. 2004 ; VERTIDOS AG-EBS-250 • >200 sPc; 42° 50.507′N , 009° 25.773′W to 42° 48.810′N, 009° 25.198′W; 151– 148 m ; 17–28 SeP. 2004 ; VERTIDOS CA-EBS-150 • 30 sPc; 42° 15.780′N , 009° 10.500′W to 42° 17.100′N, 009° 10.630′W; 149–150 m ; 17–28 SeP. 2004 ; VERTIDOS CH-EBS-150 • 8 sh; 43° 29.13′N , 008° 43.233′W to 43° 29.596′N, 008° 42.838′W; 149–151 m ; 15–24 Jul. 2008 ; A SELVA DRN-22 • 14 spc; 43° 31.727′N , 008° 52.374′W to 43° 32.678′N, 008° 52.687′W; 173–179 m ; 15–24 Jul. 2008 ; A SELVA DRN-23 • 12 sPc; 44° 06.496′N , 008° 23.522′W to 44° 07.16′N, 008° 23.201′W; 337– 324 m ; 15–24 Jul. 2008 ; A SELVA AT-13 • 1 spc + 1 sh; 44° 08.516′N , 008° 40.788′W to 44° 09.688′N, 008° 39.721′W; 436– 428 m ; 15–24 Jul. 2008 ; A SELVA EBS-11 • 14 sPc; 44° 05.687′N , 008° 24.558′W to 44° 07.253′N, 008° 24.302′W; 337– 330 m ; 15–24 Jul. 2008 ; A SELVA EBS-13 • 15 sPc; 44° 00.806′N , 008° 33.098′W to 44° 01.967′N, 008° 32.57′W; 346– 344 m ; 15–24 Jul. 2008 ;A SELVA EBS-14 • 17 sPc + 10 sh; 43° 58.84′N , 008° 15.625′W to 43° 59.621′N, 008° 14.285′W; 233–237 m ; 15–24 Jul. 2008 ; A SELVA EBS-20 • > 50 sPc; 43° 31.687′N , 008° 52.188′W to 43° 33.149′N, 008° 53.142′W; 171–182 m ; 15–24 Jul. 2008 ; A SELVA EBS-23 • 5 sPc; 43° 48.252′N , 008° 51.427′W to 43° 49.707′N, 008° 51.164′W; 575–584 m ; 15–24 Jul. 2008 ; A SELVA EBS-30-1 • 4 sPc + 3 sh; 43° 33.20′N , 008° 58.78′W to 43° 34.57′N, 008° 56.90′W; 230–236 m ; 15–30 SeP. 2008 ; DIVA-Artabria II EBS-01 • 1 sh; 43° 34.86′N , 009° 21.95′W to 43° 36.44′N, 009° 20.71′W; 1010–1112 m ; 15–30 SeP. 2008 ; DIVA-Artabria II EBS-04 • 2 sPc; 43° 19.92′N , 009° 38.13′W to 43° 21.25′N, 009° 37.11′W; 1267–1286 m ; 15–30 SeP. 2008 ; DIVA-Artabria II EBS-15 • 1 sh; 42° 31.58′N , 009° 40.14′W to 42° 33.87′N, 009° 38.75′W; 2030– 1886 m ; 15–30 SeP. 2008 ; DIVA-Artabria II EBS-30 . FIGURE 5. a–d Alvania punctura ( Montagu, 1803 ) . a Shell (1.8 mm) (DA-I(02): DRN-1000); b Young shell (1.2 mm) (VE: AG-EBS-150); c Lateral view of the protoconch; d Details of the microsculpture of the teleoconch. e–g Alvania testae (Aradas & Maggiore, 1844) (VE: AG-EBS-150). e Shell (2.2 mm); f Lateral view of the protoconch; g Detail of the microsculpture. Remarks. Very common species that extends from Norway to off western Morocco and throughout the Mediterranean, in a wide bathymetric range 30–1000 m , occasionally deeper ( Oliverio et al . 1992 ; Bouchet & Warén 1993 ; Warén 1996 ; Cachia et al . 1996 ; Giribet & Peñas 1997 ; Gofas & Oliver 2011 ; Negri & Corselli 2016 ; Bitlis & Öztürk 2017 ). The shell is characterized by the opisthocline orientation of the peristome ( Fig. 5e ). Protoconch sculptured by an irregular network ( Figs. 5f–g ). It was the most abundant species in studied the samples along with Obtusella macilenta , predominantly between 100 and 400 m in muddy sand bottoms. Its putative planktrotrophic larval development (deduced from its protoconch) enables a large dispersal capacity.