Amphibians of Togo: taxonomy, distribution and conservation status Author Segniagbeto, Gabriel Hoinsoudé Laboratory of Ecology and Ecotoxicology, Faculty of Sciences, University of Lomé, BP 6057 Lomé, Togo, BP 1515, Lomé (Togo) and Institute for Development, Ecology, Conservation and Cooperation, Rome (Italy) gsegniagbeto @ gmail. com (corresponding author) Author Mark-Oliver, Annemarie Ohler Author Luca, Mark-Oliver Rödel Author Alain, Luca Luiselli Author Dubois, Alain Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE-PSL, Université des Antilles, 57 rue Cuvier, CP 30, F- 75005, Paris (France) alain. dubois @ mnhn. fr dubois@mnhn.fr text Zoosystema 2024 2024-10-08 46 25 631 670 https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a25.pdf journal article 10.5252/zoosystema2024v46a25 1638-9387 13907152 D4AC1F89-AC34-43C4-9761-3F2015A02265 Ptychadena pumilio (Boulenger, 1920) ( Fig. 13A ) MATERIAL EXAMINED . — Togo • 7 ♂ , 10 ♀ ; Kanté; MNHN- RA-1996.8861-8863, MNHN-RA-2006.2219, MNHN- RA-2006.2221,MNHN-RA-2006.2223-2229,MNHN-RA-1996.8322, MNHN-RA-1996.8323, MNHN-RA-1996.8324, MNHN- RA-1996.8860 • 4 ♂ , 4 ♀ ; Mango; MNHN-RA-1996.8330-8341 • 7 ♂ ; Oga; MNHN-RA-2008.0154-0157. DESCRIPTION . — Small frog (SVL 21.3-29.8 mm ♂ , 24.4-37.1 mm ♀ ), with fairly slender body. Snout pointed, protruding above the mouth slit. Head slightly longer than broad (HW 27-34% SVL; HL 37- 45% SVL). Tympanum distinct (TYD 4-9% SVL). Hind legs long (TL 51-67% SVL). Webbing small, leaving some phalanges free; webbing formula: I 1 – 2 II 1 – 2 ½ III 1 ½ – 3 IV 3 – 1 V. Tip of toes pointed. Back marked by four pairs of glandular folds, the inner most pair starting from the back of the eye, running along the medial dorsal line and ending at the vent, the third pair being the shortest and in a median position with respect to the first two, the fourth, the most outer, being dorsolateral; distinct and very short supratympanic fold; smooth skin on the back and more granular on the flanks. COLOURATION . — The colouration is olive-gray or brown on the back, with a whitish, orange or green vertebral line. Some darker spots can be seen on the back. The dorsolateral folds are light gray. From the tip of the snout to the eye, the canthus is marked by a dark band. The dorsal parts of the forearm, thigh and tibia are crossed out by darker bands. The belly is white. SEXUAL DIMORPHISM . — The males are distinguished by a pair of vocal sacs, opening on either side of the throat, parallel to the lower jaw, and nuptial pads on first to third fingers. HABITATS AND DISTRIBUTION . — Ptychadena pumilio is essentially a savannah species. It is distributed in ecological zones I, II, III and IV. Its frequency in the ecological zone IV becomes more and more important because of the savannisation of the forest zone. It has been reported in Togo by Bourgat (1979) , Bourgat et al. (1996) and Segniagbeto et al. (2007 , 2022 ).