A revision of the genus Rudolfina Roháček (Sphaeroceridae: Limosininae) Author Paiero, Steven Mark Author Marshall, Stephen A. text European Journal of Taxonomy 2020 2020-01-16 593 1 48 journal article 24112 10.5852/ejt.2020.593 16dfaaea-fa3b-4dda-a047-13359b05bfa1 3659777 B08C59C9-6F91-4F3B-B595-5C6E7E6710D0 Rudolfina megepandria sp. nov. urn:lsid:zoobank.org:act: 16E7249F-BC7E-4CD0-AFCB-60A5C98CFB43 Figs 4 , 13 , 21B Etymology The specific name, which has been a manuscript name since Marshall (1982) , refers to the unusually large epandrium. Material examined Holotype MEXICO • ♂ ; Jalisco , 18 mi. W of Atenquique ; 9300 ft a.s.l. ; A. Newton leg.; fir forest, dung; FMNH debu01086084. Fig. 13. Rudolfina megepandria sp. nov. , male terminalia (debu01086085). A . Epandrium, surstylus and cercus, lateral view. B . Same as preceding, caudal view. C . Sternites 4–7, ventral view. D . Phallus and postgonite, dorsal view. E . Phallus, postgonite and phallapodeme, dorsolateral view. F . Same as preceding, lateral view. Scale bars: A–B = 0.20 mm; C = 0.05 mm; D–F = 0.05 mm. Paratypes MEXICO – Jalisco • 1 ♂ , 1 ♀ ; same collection data as for holotype; DEBU . – Oaxaca • 1 ♀ ; Yuvila Rd ; 9400 ft a.s.l. ; 9–19 Aug. 1973 ; A. Newton leg.; mesic oak, carrion; FMNH . Description BODY. Length: 1.9–2.2 mm . Eye height 2.5 × genal height. Head with 3 interfrontal setae and row of 4–5 small setulae on inner margin of orbital plate. Gena with 2 strong setae and 7–8 smaller setulae. Acrostichal setulae in 4–6 rows. Costagial seta extending to subcostal break. Second costal sector 0.75– 0.8× third costal sector. Length of M 1 between crossveins dm-cu and r-m– 3.0× dm-cu; CuA 1 stub vein ~10.0 × M 1 stub vein (M 1 stub vein extremely reduced). Male mid femur with 14–15 strong setae in ventrobasal cluster. Male mid tibia with ventral comb of 4–5 robust, short setae on apical ½. Mid tibia without midventral seta (both sexes). MALE ABDOMEN ( Fig. 13 ). Sternite 5 posterior margin with 2 small nipple-like teeth on each side of small emargination; emargination ~1/15 width and ~1/9 length of sternite 5. Transverse part of sternite 6 arcuate. Epandrium swollen, broader than preceding abdominal segments; subanal plate broad, complete. Surstylus with posterior lobe elongate, length ~4.0× basal width, with a hirsute medial lateral lobe; apex truncate. Cercus elongate-conical, projecting posteriorly. Postgonite elongate, narrowly rounded distally and narrow basally. Dorsal sclerite of distiphallus ( Fig. 13 D–F) without dorsal swellings and not extending beyond apex of acrophallus. FEMALE ABDOMEN ( Fig. 4 ). Tergite 7 with posterior margin entire. Medial part of tergite 8 elongate, length ~2.0–3.0 × width, with posterior margin emarginate. Epiproct broadly rounded anteriorly. Cercus ~0.75× length of epiproct; fused anterolaterally to epiproct. Sternite 7 with posterior margin broadly rounded. Spermathecae cup-shaped; paired spermathecae with stems ~1.5× width of spermatheca. Remarks Rudolfina megepandria sp. nov. is diagnosed by its swollen male epandrium and distinctively shaped sternite 5. It is most closely related to R. bucki sp. nov. , which has a less pronounced epandrium and differs in details of the surstylus and cercus.