Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia Author Grisales, Diana Author Wolff, Marta Author De, Claudio J. B. text Zootaxa 2012 3591 1 46 journal article 10.5281/zenodo.213946 2a54e127-4d40-404e-ab7f-5b48f6e37e83 1175-5326 213946 Fannia grandis Malloch ( Figs. 6, 7 , 26 , 44 , 62 , 80 , 98 , 116 ) Fannia grandis Malloch 1912 : 3 . Holotype male, deposited in the National Museum of Natural History (USNM). Type-locality: Panama , Puerto Bello. Diagnosis. these characters apply only to the male sex. Eyes with sparse setulae, hind femur with ventral well developed pre-apical protuberance and a large tuft of densely packed setae, forming a strong hook. Redescription . Male: body length: 7.2 mm; wing length: 5.25 mm. Head : eye with sparse setulae, short setae visible under certain angles. Frontal vitta velvety brown with orange areas, posteriorly black-greyish and pollinose. Interocular space 0.16 mm. Fr 14. Ocellar triangle greyish pollinose; oc proclinate and developed, 5 less developed setulae in the area. Poc divergent. Fronto-orbital plate and parafacial greyish pollinose. Parafacial bare. Gena grey. Lunule yellow. Scape yellow, 3–5 short divergent setae. Pedicel yellow, with dorsal setae short ( Fig. 6 ). Postpedicel golden-yellow pollinose, 2 times the length of pedicel. Arista brown, base and basal fourth yellow, pubescent. Palpus yellow and filiform. Thorax : grey, golden-brownish pollinose; scutum with 3 ample brown vittae, median vitta wide and irregular, ending at base of scutellum, the two lateral vittae wide, reaching the postpronotal, pre-sutural and intra-alar areas and ending on the last pair of postsutural dc ; in dorsal view vitta expanded to supra-alar area; in lateral view only a brown mark is seen on supra-alar area ( Fig. 7 ). Scutellum brown on basal half. Acr 2–3:3–4, not arranged in rows; 2 setae shorter than pre-scutellar setae near base of scutellum. Dc 2:3, arrangement of postsutural dorsocentral similar to the base of a cone.Postpronotum with 2 developed and 2 weak setae. Two short pra , larger seta closer to supra-alar seta. Proepimeral area with few weak ground setulae. Pre-basal pre-scutellar seta absent, area with short ground setulae. Subapical scutellar setae 1 pair, on the same row as discal scutellar setae. Lateral scutellar setae present. Wing : yellowish. Calypters yellowish with margin darker, lower calypter triangular with weak, brownish apical mark. Haltere yellowish with stem whitish and base yellowish. Legs : brown with coxae yellowish-brown and silver-golden pollinose; trochanter yellow; femur light-brown with base and apex yellow; tibia with basal third and apex yellow; tarsi dark brown and pulvillus yellowish ( Fig. 6 ). Fore femur with 1 row of d ; ventral surface almost bare; 1 row of long and strong pv on apical half; 1 row of pd ; 1 row of p, longer and with curved apices on apical third. Fore tibia with 1 strong pre-apical d ; 1 apical pv . Fore tarsus with basal v on first tarsomere, with 1–2 long and weak setae; tarsomeres, particularly the two apical most, slightly flattened and widened with respect to the remaining tarsomeres. Mid femur constricted on pre-apical ventral surface; 1 row of ad ending in 1 long and curved pre-apical; 1 row of short av longer and sparse on basal third and decrease in length towards apex; ventral surface with strong and developed setae on basal half and on apical half, on apical third with hooked apices and running towards posterior surface, forming a ctenidium; 1 row of long p ending on posterodorsal surface, with hooked apices on basal half and 5 strong and straight setae on apical half. Mid tibia on ventral surface strongly constricted, on apical half densely setulose and with 1 apical long and strong seta; 1 ad on edge of apical third; 1–2 short apical a ; 1 long and strong apical and 1 short and weak av ; 1 short p on edge of apical third and 1 pre-apical; 1 pre-apical d . Mid tarsus with first tarsomere 2 times the length of first tarsomere of fore and hind legs. Hind coxa bare on posterior margin. Hind femur lightly curved; ventral surface with pre-apical conspicuous protuberance, where a strong tuft of long and strong setae forms a hook ( Figs. 26 , 44 ); 1 row of ad running towards dorsal surface, increasing in length towards apex and ending in 7 developed setae; anterior surface with setae on basal half; 1 row of long and strong av on apical half up to pre-apical protuberance, 1 long and strong seta on apical third and 3 shorter apical ( Fig. 26 ); posterodorsal and posterior surfaces densely setulose, with long and weak setae with hooked apices, more ventral setae extending up to protuberance ( Fig. 44 ). Hind tibia with dorsal surface curved, 1 strongly developed median and 1 shorter pre-apical d ; 2 median ad , 1 developed on edge of apical third and 1 shorter, pre-apical; 1 row of short and strong a on apical half, unordered, and 2 short pre-apical; 5–6 av on apical half, 1 strong apical. Abdomen : syntergite 1+2 with base brown; syntergite 1+2, tergites 3–4 translucent-yellow or darker, with median vitta brown, expanded on apical margin of each tergite without reaching lateral region; tergite 5 grey pollinose and with median vitta brown, expanded to apical margin ( Fig. 7 ). Sternite 1 bare. Sternite 5 as in Fig. 62 . Terminalia ( Figs. 80 , 98 ): epandrium wider than long, with developed setae; cercal plate enlarged with weak setae and two apical, long projections; surstylus strongly articulated with epandrium, straight and pointed apically, with short and weak setae and apex folded; bacilliform process spiralled; hypandrium and associated structures as in Fig. 116 . Female : not found. Biology. label data of examined material from Colombia indicate that this species is associated with decomposing organic matter in forest areas. Comments. Fannia grandis is a new record for Colombia . Until now the species was known only from Panama , collected by the biological entomological survey of the Smithsonian Institution in the Panama Canal in 1911 ( Malloch 1912 ). According to Albuquerque et al. (1981) the species belongs to the grandis group. However, the phylogenetic placement of F. grandis is in the following clade: [ F. h e r m a n i [ F. grandis [ F. ro i g i + F. losgateados ]]], sister-group of the anthracina group ( Domínguez & Roig-Juñent 2008 ). Type-material (not dissected): Paratypes (USNM). PortoBello\ Pan [ Panama ] Mar 13. 11 \ August Busck. Paratype \ No 14912\ U.S. N.M ( 1 male ); PortoBello\ Pan [ Panama ] Feb 25.11\ August Busck Collector. Paratype \ No 14912\ U.S. N.M ( 1 male ); PortoBello\ Pan [ Panama ]\ Feb 23 \ A Busck Coll. Paratype \ No 14912\ U.S. N.M. ( 1 male ). Material examined. COLOMBIA . Antioquia: 1 male San Roque, Cto. San Jos del Nus, Corpoica, VSR liver, Borde de Bosque, N. Uribe, May 0 1, 2007, CEUA 42629 ( CEUA ). Distribution. Panama ( Malloch 1912 ; de Carvalho et al . 2003 ). Colombia : department of Antioquia (Andean region, Central Cordillera).