Revision of Silhouettanus with description of nine new species (Hemiptera: Heteroptera: Schizopteridae)
Author
Hill, Lionel
text
Zootaxa
2014
3815
3
353
385
journal article
45480
10.11646/zootaxa.3815.3.3
d77979aa-7cff-4879-a358-73f3c2f97557
1175-5326
226610
4A7AAB28-5455-4D3C-B642-463376AC6A94
Silhouettanus bamaganus
sp. n.
(
Figs 2–3
, dimensions in
Table 1
)
Types
material
.
Holotype
: T189823, mac. ♂, Bamaga, NQ,
19–29 Jun 2009
, J. Bond and J. Sailor, ex
Culicoides
trap in cattle paddock, dissected on seven slides in QM.
Paratypes
: T189824-5, 1 mac. ♂, as for
holotype
, whole in vial in QM; 1 mac. ♂, NEQ, Bramwell Station via Cairns,
18–20 Nov 2008
, T. Kerlin and B. Cookson, ex
Culicoides
trap, left fore wing on slide, remainder undissected in vial in QM.
Description
. Brown with two cream bands, that is clavus and adjoining corium and distal membrane cream. L4 relatively long, ratio of lengths of labial segments 35:30:13:42; costal lobe wide; divergence of R+M from Sc midway along lobe, distal to the proximal m-cu cross vein therefore subcostal cell short (R+M is parallel and contiguous with C+Sc proximally); no emargination of membrane at apex of costal lobe; cell R <discal cell, divergence of M from R distal to basal cell, more than half area of cell R is distal to apex of discal cell; 1AN joins Cu opposite tornus; vein setae medium. Right side of T8 produced as elongate, curved strut widening distally and embracing right anterodorsal part of genital capsule, with spiracle apically; left side T8 shortly and broadly produced with spiracle apically; left anophoric process is a dark, broad, apically acute process that half-sheaths a spinous, parallel branch; right anophoric process is a long, curved strut embracing genital capsule and accommodated by its contours; left paramere short, bulbous base tapering to an acute apical lobe; right paramere elongate, curved, with bilobate apex of which one lobe is slender, curved and digitate; one conjunctival sclerite bearing a large crescentic process with acute apex (major process,
Fig. 3C, C
1
); second conjunctival sclerite bearing a short curved spine (minor process,
Fig. 3C, C
2
); vesica with one sclerotised loop.
FIGURE 1.
A–E,
Silhouettanus alboclavatus
male; A, fore wing, dorsal; B, wing coupling, dorsal through wing; C, labium and antennal base; D, pterosternum; E, T8 and genital capsule, dorsal (lap and rap in Fig. 1E should be transposed). Abbreviations: ac, anal cell; bc, basal cell; C1, major conjunctival process; C2, minor conjunctival process; dc, discal cell; gc, genital capsule; lap, left anophoric process; lp, left paramere; m, metasternal process; me, metendosternite; ms, mesosternal sulcus; rap, right anophoric process; rc, cell R; rp, right paramere; scc, subcostal cell; t, tornus; tc, trapezoidal cell; v, vesica; II, midcoxa.
FIGURE 2.
A–G,
Silhouettanus bamaganus
male; A, anteclypeus, antennal base, labrum and stylets, dorsal; B, labium, lateral; C, right fore leg; D, fore claw and parempodium; E, right midleg; F, right hind leg; G, fore wing, dorsal. Abbreviations: AC, anteclypeus; bc, basal cell; dc, discal cell; gc, genital capsule; LR, labrum; rc, cell R; scc, subcostal cell; tc, trapezoidal cell.
FIGURE 3.
A–C,
Silhouettanus bamaganus
male; A, T8, anophore and genital capsule, dorsal; B, parameres and aedeagus, sketch during dissection; C, parameres and aedeagus. Abbreviations: a, anophore; bp, basal plate; C1, major conjunctival process; C2, minor conjunctival process; gc, genital capsule; lap, left anophoric process; lp, left paramere; rap, right anophoric process; rp, right paramere; scc, subcostal cell; t, tornus; tc, trapezoidal cell; v, vesica.
Notes
. The large crescentic conjunctival process is similar in size to that of
S. alboclavatus
. The long right process of T8 was not observed in other
Silhouettanus
species described here and resembles that of
Pinochius josifovi
Redei, 2008
(
Rédei 2008: fig 13
). Its presence means that two not one curved struts embrace the genital capsule laterally on the right in this new species. The contiguity of R+M to C+Sc for half the length of the costal lobe may be apomorphic. Undescribed, macropterous females are known from Moa and Saibai Islands in Torres Strait not far from Bamaga at the tip of Cape York Peninsula. Another species of
Silhouettanus
, not described here, also occurs at Bamaga. It is notable for a medial pit on the pronotal sulcus of males. Such a pit was not observed in any other species described here but is probably analogous to pits or sensory areas that occur in the males of several other schizopterid genera variously on the anteclypeus, postclypeus, frons, vertex, pronotum and clavus. The male of this undescribed species has R+M separating basally from C+Sc, two pale bands across the fore wing, cell R subequal to the discal cell, a vesica with one loop and an outwardly recurved apex on the spinous, curved major conjunctival process.
Etymology
. Name refers to the
type
locality near the tip of Cape York Peninsula; adjective.