Synopsis of the Grenadier Fishes (Gadiformes; Teleostei) of Taiwan
Author
Iwamoto, Tomio
Section of Ichthyology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA;
Author
Nakayama, Naohide
Laboratory of Marine Biology, Faculty of Science, Kochi University, 2 - 5 - 1 Akebono-cho, Kochi, 780 - 8073, Japan;
Author
Shao, Kwang-Tsao
Research Center for Biodiversity, Academia Sinica No. 128, Sec. 2, Academia Road, Nankang, Taipei 115, Taiwan;
Author
Table, Hsuan-Ching Ho
text
Proceedings of the California Academy of Sciences
2015
2015-04-15
62
3
31
126
journal article
299670
10.5281/zenodo.11512126
522b8de7-880d-40ca-803e-aa5a3a05c5f6
0068-547X
11512126
Nezumia proxima
(Smith and
Radcliffe, 1912
)
Figure 19
.
Macrourus proximus
Smith and Radcliffe
in
Radcliffe, 1912:119–120
, pl. 26, fig. 2 (
holotype
,
292 mm
TL,
USNM 72936
;
Sogod Bay
,
Leyte
I.,
Philippines
,
10°12ʹ00ʺN
,
125°04ʹ10ʺE
, 502 fm [
918 m
];
paratypes
,
USNM 135338–39
).
Macrourus nasutus
[non
Coryphaenoides nasutus
Günther, 1877
]:
Jordan
and Gilbert, 1904:618 (spec. from off Izu,
Japan
).
Lionurus proximus
: Gilbert and Hubbs, 1916:201–202 (
11 spec.
[excluding
3 types
];
East
China
Sea
,
Pacific
coast of s.
Japan
, 361–544 fm [
660–995 m
]).
Lionurus
(subgenus
Nezumia
)
proximus
: Gilbert and Hubbs, 1920:554 (
3 spec.
[including
holotype
],
Philippines
).
Lionurus abei
Matsubara, 1943:146–147
, fig. 7, 8 (
holotype
,
160.7 mm
TL, cat. no. 4909, Kumano-nada,
Japan
; “
Sigenkagaku Fish
Coll. No.
7”).
Nezumia proximus
:
Okamura, 1970:94–98
, pl. XXI, text-fig. 41(
17 spec.
,
174–370 mm
TL; e. coast s.
Japan
, East
China
Sea;
420–910 m
; synonymized
Lionurus spinosus
Gilbert and Hubbs, 1916
with
N. proxima
).
Nezumia proxima
: Okamura
in
Okamura et al., 1982:163
, 351, fig. 98 (p. 162) (
1 spec.
,
160 mm
TL; Kyushu-Palau Ridge).— Sawada
in
Amaoka et al., 1983:107
, 192, fig. 58 (p. 106) (
4 spec.
, 269.6–346.
4 mm
TL; Ibaragi [central Honshu, ca.
37°N
],
Japan
,
1100 m
).— Okamura
in
Masuda et al., 1984:95
, pl. 81–I (compiled).— Okamura
in
Okamura and Kitajima, 1984:217, 362, fig. 152 (p. 216) (
8 spec.
,
186–350 mm
TL;
Okinawa
Trough [East
China
Sea],
820–932 m
).—
Iwamoto, 1990:281–282
, fig. 645–646 (compiled).—
Nakabo, 2002:424
(compiled).—
Shao et al., 2008
: table 2 (
2 spec.
,
Taiwan
[
ET
,
SCS
],
488–1098 m
; first record from Taiwan).
MATERIAL
EXAMINED
(
2 spec.
).—
NET
:
ASIZP 65632
(1, 377+ TL)
, CP 214, 488–
1027 m
.
SCS
:
ASIZP 66187
(1, 32 HL, 163+ TL)
; CD 322,
1098 m
.
SWT
:
ASIZP 65529
(1, 282+ TL)
, CD 142, 227–
335 m
.
DISTINGUISHING
FEATURES
.— 1D II,9–11; P i17–i21;
V
9, rarely 10; inner GR-I 9–11 total; scale rows below midbase 1D 5–7.5, below 2D 7–8.5; pyl. caeca 24–31. Snout length 23–31% HL; interorbital 18–25%; orbit 28–33%; suborbital 14–17%; upper jaw 31–34%; barbel 15–23%. Body relatively slender, compressed; head in TL about 6–7 times; snout conical, protruding well beyond mouth, with stout tubercles at tip and lateral angles; suborbital ridge sharp and beset with stout modified scales; underside of head mostly scaled except for broad naked median swath on underside of snout; body scales densely covered with recumbent narrowly triangular to lanceolate spinules. Long spinous ray of 1D about 70–100% HL, serrations strong, sharp, widely spaced. Anterior dermal window of light organ small, situated between inner V-fin bases; anus about midway between
V
and A. Color overall dark brown, abdominal region bluish to black; gill cover, gill membranes black; mouth dusky; fins dark dusky to black. Attains at least
380 mm
TL.
DISTRIBUTION
.— Broadly distributed from east-central
Japan
and East
China
Sea to South
China
Sea off
Taiwan
and the
Philippines
; recorded depth range
355–1100 m
. The sw.
Taiwan
specimen was taken in
227–335 m
; the ne.
Taiwan
specimen in
488–1027 m
, and the
SCS
specimen in
1098 m
.
REMARKS
.— The
133 mm
specimen listed in the table in Gilbert and Hubbs (1916:202) as
N. proxima
from
Albatross
sta. 4918 is deposited at
CAS
[cat. no.
CAS-SU
22941]; we have determined that it represents
N. spinosa
, a species that is closely similar to
N. proxima
and with which it has been mistaken in the past. However, the underside of head of
N. spinosa
is almost wholly naked, compared with only the median portion of snout in front of the mouth naked in
N. proxima
. The spinous 1D ray is also longer in
N. spinosa
, the scale spinules needle-like and not flattened (viz., not lanceolate or spear-shaped), and the outer premaxillary dentition is larger than in
N. proxima
.