Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species
Author
Gunter, Nicole L.
Author
Weir, Thomas A.
text
Zootaxa
2017
4290
2
201
243
journal article
32718
10.11646/zootaxa.4290.2.1
43ebda6c-201a-46af-8b54-e7980b8d79e1
1175-5326
828965
52D6E3C3-BA0A-49C7-865C-00F3F5E5C5D4
Monteithocanthon
Gunter & Weir
,
new genus
Type species.
Lepanus arator
(
Matthews, 1974
)
Etymology.
Named in honour of Geoff Monteith for his contribution to the taxonomy of Australian dung beetles.
Diagnosis.
Small, total length
2.1–3.2 mm
. Elytra with 8 striae, 7 on disc and one on edge of epipleura (
Fig. 7
A). Seventh elytral striae obsolete (
Fig. 7
A). Pygidium with transverse groove at the base not reaching basal angle (
Fig. 9
). Head without basal carina. Dorsal parts of eyes narrow, 2–5 facets wide; narrowest at base of head; eye canthus dividing or nearly dividing eye. Antennae with 3-antennomere club with club antennomeres of approximately equal width. Protibiae of male with comb of stiff setae and ventral brush of setae at inner apex; apical spur small, inconspicuous (
Fig. 7
C–D). Protibiae with 3 teeth on outer edge, not equidistant with apical 2 slightly closer together; serration between teeth and proximal to them (
Fig. 7
D–F). Each tooth has 1 group of setae dorsally (
Fig. 7
D). Distal edge of protibiae straight and trochantofemoral anterior pit absent (
Fig. 7
B). Trochanter excavated dorsally for reception of inner apical edge of protibia. Protarsi with 5 tarsomeres, tarsomere 5 the longest. Metatarsomere 1 wider than 2–5, metatarsomere 2 equal to tarsomere
5 in
length (
Fig. 7
J). Male mesotibiae with dense setae on inner edge (
Fig.
7
I). Claws toothed (
Fig. 7
C). Male metatibiae generally bent with an extension but almost straight in
M. peckorum
(
Fig. 8
). Mesometasternal suture procurved to strongly procurved. Mesepimera impunctate. Aedeagus with parameres asymmetrical, right paramere apically bent downwards and more-or-less expanded apically as in
Fig. 9
. Hind wings fully developed, reduced, or absent.
Remarks.
Matthews (1974)
retained the genus
Lepanus
for practical purposes noting that adhering to a strict generic concept in such a morphologically diverse genus would lead to fragmentation into several small, difficult groups. All species of
Monteithocanthon
share many morphological affinities with
Lepanus
so it is unsurprising that
M. arator
and
M. glaber
were originally described within the genus
Lepanus
. A new genus is established here primarily on the basis of genetic results, with the two
Monteithocanthon
species sequenced to date always distinct from the clade containing the majority of
Lepanus
species. The phylogenetic grouping and the establishment of a new genus are supported by morphology. The six species assigned here to
Monteithocanthon
share a number of morphological features in combination, although many of these diagnostic characters are also individually found in other Australian ‘deltochiline-like’ genera and species.
Distribution.
The genus is known only from
Australia
. Four species occur in northern
Queensland
primarily in the Wet Tropics, one species is distributed at high altitude in southeastern
Queensland
, and one species in southeastern
Western
Australia
(
Fig. 5
).