Review and reclassification of the Old World genus Physopelta (Hemiptera: Heteroptera: Largidae) Author Stehlík, Jaroslav L. Department of Entomology, Moravian Museum, Hviezdoslavova 29 a, CZ- 627 00 Brno – Slatina, Czech Republic text Acta Entomologica Musei Nationalis Pragae 2013 2013-11-15 53 2 505 584 journal article 2413 10.5281/zenodo.5739959 956fa4e4-8b18-4c70-94b2-5722d5348180 0374-1036 5739959 8D64B275-C6AF-411E-B79D-A3EBEB3D093FD Physopelta ( Neophysopelta ) gutta gutta ( Burmeister, 1834 ) ( Figs 17–34 , 98 ) Lygaeus ( Pyrrhocoris Fall., Platynotus Schill.) gutta Burmeister, 1834: 300–301 , pl. XLI: fig. 10. SYNTYPE (s): Philippines , Manila env. (ZMHB, 1 ♂). Pyrrhocoris gutta : BURMEISTER (1835) : 285 (new combination, diagnosis, distribution). Physopelta bimaculata Stål, 1855:186 (description, distribution). SYNTYPE (s): Indonesia , Java (NHRS, G. Lindberg, pers. comm.). Synonymized by STÅL (1861: 195) . Physopelta gutta : STÅL (1861) : 195 (new synonymy, list); STÅL (1863) : 391 (catalogue, distribution); STÅL (1870) : 99–100 (key to species, catalogue, distribution); STÅL (1871) : 665 (list of species); WALKER (1873) : 18, 19 (key, catalogue, distribution); SCOTT (1874) : 291 (distribution); DISTANT (1879a) : 127 (distribution); DISTANT (1879b) : 37 (distribution); DISTANT (1883) : 417 (distribution); LETHIERRY (1888) : 463 (distribution); KIRBY (1891) : 105 (distribution); LETHIERRY & SEVERIN (1894) : 242 (catalogue, distribution); UHLER (1897) : 265 (variability, distribution); BREDDIN (1900) : 161 (distribution; not distinguishing Ph. g. famelica ); BREDDIN (1901b) : 139 (distribution); DISTANT (1903a) : 97 (redescription, figures, distribution); DISTANT (1903b) : 250 (catalogue, distribution); MAXWELL- LEFROY (1909): 325 (distribution); ESAKI (1926) : 157 (distribution); TAEUBER (1927) : 174–177 (redescription, variability, distribution; differences from Ph. lisae and Ph. roseni , respectivelly); BLÖTE (1931) : 99–100 (distribution); SCHMIDT (1931) : 46 (distribution); LIU (1981) : 222, 224 , pl. 28 (key, redescription, figure); AHMAD & ABBAS (1987) : 137–138, 141–142 (redescription, figures, host plant, distribution); AHMAD & PERVEEN (1986) : 162, 164–166 (morphology of abdomen and genitalia); AHMAD & PERVEEN (1989) : 49–50 (cephalic morphology); ZAMAL & CHOPRA (1990) : 4 –5 (redescription, figures, distribution); LEE & KWON (1991) : 50 (distribution); TOMOKUNI (1993) : 59, 197 (redescription, habitus, larva, egg, biology); SEN et al. (1998) : 331, 333–334, 339–340 (checklist, key, diagnosis, distribution); HUA (2000) :187 (check-list, host plants, distribution); KWON et al. (2001) : 301–302 (check-list, host plants, distribution); TOMOKUNI (2006) : 352 (distribution); SAHA & BAL (2007) :322–324 (checklist, key, diagnosis, distribution); CHANDRA et al. (2012) :70–71 (figure, distribution); CHANDRA & KUSHWAHA (2012) : 253 (distribution); CHANDRA & KUSHWAHA (2013) : 678 (distribution); SINGH & BANYAL (2013) : 1056 (distribution). Physopelta gutta gutta : HUSSEY (1929) : 30 –31 (catalogue, distribution); ESAKI (1952) : 230 (redescription, habitus, distribution); STEHLÍK (1965a) : 287 (identification of gutta and famelica , distribution); MANNA et al. (1985) : 621–630 (chromosomes, figures, distribution); CHAKRABARTY et al. (1994) : 34–35 (differential diagnosis, distribution); KERZHNER (2001) : 246 (catalogue, distribution); CASSIS & GROSS (2002) : 626–627 (catalogue, distribution – mixed with gutta famelica ); KOHNO et al. (2002) : 393 (distribution); STEHLÍK & JINDRA (2003) : 7 (distribution); STEHLÍK (2004) : 2–3 (description of 5th larval instar, description); STEHLÍK (2005a) : 144 (distribution); STEHLÍK (2007a) : 117–118, 127 (habitus, variability, identification and distribution of gutta and famelica ); RÉDEI et al. (2009) :8, 12–15, 48 (redescription, key, figures, distribution); KMENT & VILÍMOVÁ (2010) :4, 11–12 (morphology of external scent efferent system). Physopelta ( Physopelta ) gutta gutta : VOIGT (2006) : 224 (key, distribution – mixed with gutta famelica ). Physopelta ( Physopelta ) gutta famelica (misidentification): VOIGT (2006) : 224 (record from China). Physopelta guttata (incorrect subsequent spelling): YANG (2003) : 200–201 (morphology, figures). Material examined. JAPAN : KYUSHU: Korasan, Chikugo, 1.viii.1955 , 1 ♂, 25.ix.1955 , 1 ♀ , S. Miyamoto lgt. & det. (NMPC). – RYUKYUS: OKINAWA ISLAND: W slope of Katsudake Motobu Pen., 28.ix.1945 , 1 ♂, F. G. Werner lgt. (AMNH); ditto, Iwa, 28.vii.1945 , 1 ♂ 1 ♀ , F. G. Werner lgt. (AMNH). ZAMANI- SHIMA: Ketto, xi.1945 , 1 ♀ , Wh. Nutting lgt. (AMNH). – CHINA : GUIZHOU : Kouy-Tchéou [= Guizhou], Rég. de Pin-Fa, 1909, 1 ♂, Pére Cavalerie lgt., P. Kment det. (MNHN). – HAINAN : Bawangling Mts., Baote env., 19°05.3′N 109°07.4′E , 415–800 m a.s.l., 7.–8.v.2011 , 1 ♀ , M. Fikáček, V. Kubeček & L. Li lgt. (NMPC). – TAIWAN : MIAOLI COUNTY: hills SW Sanyi, sand-steppe, 24°22′N 120°44′E , 200–500 m a.s.l., 27.x.2008 , 1 ♂, L. Dembický lgt. (MMBC). – NANTOU COUNTY : Meifeng, 24°05′25″N 121°10′21″E , 10.–11.v.2001 , 1 ♂ 1 ♀ , K. Schönitzer lgt. (ZSMC); Puli-Yüchin, Sun Moon Lake, 16.–18.vi.1993 , 2 ♂♂, J. Dalihod lgt. (ZJPC); road No. 14, NE Puli Reyen, Shi-Reg. Meifeng, Tamblingan Lakes, Nat. Res., 24°06′N 121°10′E , 2.200 m a.s.l., 9.–11.xi.2002 , 1 ♀ , W. Schlacht lgt. (ZSMC). – NEW TAIPEI CITY : Shiuesham Mts., Fusham Botanical Garden, ca. 24°46′N 121°35′E , 650 m a.s.l., 17.–22.vi.2000 , 1 ♂ 1 ♀ , W. Schlacht lgt. (ZSMC). – TAITUNG COUNTY : 14 km W of Chihshang, 29°09′N 121°04′E , 900 m a.s.l., 16.xi.2008 , 30 ♂♂ 32 ♀♀ , L. Dembický lgt. (MMBC, NMPC). – AFGHANISTAN : KABUL : Sarobi [= Sarowbi or Surobi], 1100 m a.s.l., 16.x.1961 , 2 ♀♀ , G. Ebert lgt., L. Hoberlandt 1984 det. (NMPC). NURISTAN : Bashgultal [= Landaisin river aka Bashgul river], 1300 m a.s.l., 24.v.1953 , 1♀ , J. Klapperich lgt., L. Hoberlandt 1984 det. (NMPC). – PAKISTAN: PUNJAB: Murree Hills, Bhurban, 1,900 m a.s.l., 3.–6.v.1978 , 1♂, C. Holzschuh lgt. (EHIA); Murree, no date, 2 ♀♀ , coll. Atkinson (BMNH). – INDIA : HIMACHAL PRADESH: Simla [= Shimla], 3.–4. ix. 1975 , 2150 m a.s.l., 2 ♀♀ , A. Čejchan & L. Hoberlandt lgt. (NMPC). – WEST BENGAL : Boutan, Pedong, 1897, 2 ♀♀ , coll. Oberthür, P. Kment det. (MNHN). – NEPAL : BAGMATI ZONE: Kathmandu, British Embassy, 4500´[= 1372 m a.s.l.], 20.v.– 23.vi.1983 , 2 ♀♀ , M. J. D. Brendell lgt. (BMNH); Nagarjun, 27°45′N 85°17′E , l, 387 m a.s.l., forest, 29.vii.2000 , 1 ♀ , J. Schneider lgt. (MMBC). KARNALI ZONE : Rara Lake, 3300 m a.s.l., 6.–10.ix.1984 , 2 ♀♀ , Jt. SVC Rara Exped., M. G. Allen lgt. (BMNH). – KOSI ZONE : Kuwapani-Chichlia , 2,100–2,300 m a.s.l., 24.iv.1984 , 1 ♂ , I. Löbl & A. Smetana lgt. ( MHNG ). – BHUTAN : Bhoutan Anglais, 1900, 3 ♂♂ 2 ♀♀ , coll. Oberthür, P. Kment det. (MNHN); Maria Basti, 1897, 1 ♂ 1 ♀ , coll. Oberthür, P. Kment det. (MNHN). – SRI LANKA : Kandy env., 600 m a.s.l., 1.– 18.iv.1991 , 2 ♂♂ 1 ♀ , J. Kolibáč lgt. (ZSMC). – MYANMAR : KAYIN : Dawna [Range], 10.v.1992 , 1 ♂, S. Steinke lgt. (ZMHB). – TENASERIM : Tenass Vall, Myitta, no date, 1 ♀ , Doherty lgt., coll. Distant, P. Kment det. (BMNH). – YANGON : Rangoun [= Yangon], 1902, 1 ♂, J. Claine lgt., P. Kment det. (MNHN). – VIETNAM : AN GIANG PROVINCE: Chaadoc [= Chau Doc], 1898, 1 ♂, coll. Noualhier (MNHN). – BAC GIANG PROVINCE : Dong Lo, no date, 1 ♂, no collector, P. Kment det. (MNHN). – BAC THAI PROVINCE: Quang Chu, 25.v.1987 , 1 ♀ , Mats.-Dl.-Top. lgt. (HNHM). – CAO BANG PROVINCIE: Phia Den, 8.viii.2010 , light trap, 1 ♀ , J. Constant & P. Limbourg lgt. (ISNB); Pia Oac Mt. Nat. Res., 22°36′N 105°53′E , pine forest, 9.viii.2010 , light trap, 18♂♂ 8♀♀ , J. Constant & P. Limbourg lgt. (ISNB); ditto, 4.–6.viii.2010 , light trap, 8 ♂♂ 9 ♀♀ , J. Constant & P. Limbourg lgt. (ISNB). – HANOI PROVINCE : Hanoi, ii.1986 , 1 ♂ 1 ♀ , P. Valíček lgt. (ZJPC); Hanoi, Trau Quy – Gia Lam, I.G. 31.508, 28.xi.2004 , 1 ♀ , X. Lammens lgt. (ISNB). – HOA BINH PROVINCE : Tonkin, Hoabinh, i.1917 , 1 ♂, R. V. de Salvaza lgt. (BMNH). – LAO CAI PROVINCE : Sa Pa, v.1990 , 3 ♂♂ 2♀♀ , J. Picka lgt. (NMPC); ditto, vi.1990 , 4♂♂ 6 ♀♀ , Duong Tat Tu lgt. (NMPC); 16 km west of Sa Pa, frontier base camp, 1,800 m a.s.l., 16.iii.1989 , 3♂♂, L.Peregovits & T.Vásárhelyi lgt. (HNHM). – NGHE AN PROVINCE : forestière [= gamekeeper’s lodge] Quy Chau, forest pluv. trop. semidecid. [= tropical semideciduous rain forest], 200 m a.s.l., at light, 24.viii.1963 , 1 ♀ , T. Pocs lgt. (HNHM). – NINH BINH PROVINCE : Cuc Phuong, 3.–10.v.1966 , 1 ♂, Gy. Topál lgt. (HNHM). – QUANG TRI PROVINCE : Da Krong Nat. Park, 16°37′N 106°47′E , I.G. 31.933, 5.–10.vii.2011 , light trap, 6 ♀♀ , J. Constant & J. Bresseel lgt. (ISNB). – TUA THIEN- HUE PROVINCE: Hue, 5.iv.1983 , 1 ♀ , Macháček lgt. (NMPC). – VINH PHU PROVINCE: Me Linh Station for Biodiversity, 30.–31.v.2007 , on light, 1 ♂, P. Grotaert lgt. (ISNB); Tam Dao, 900 m a.s.l., 27.v.–2.vi.1986 , 4 ♂♂ 2 ♀♀ , J. Horák & V. Švihla lgt.; 17.–21.iii.1990 , 1 ♂ 1 ♀ , J. Horák lgt. (NMPC); ditto, 1,200 m a.s.l., l 3.x.1986 , 1 ♂, T. Vásárhelyi lgt. (HNHM); ditto, 9.v.1987 , 1 ♂, Natskási & Topál lgt. (HNHM); ditto, 200 m a.s.l., 12.x.1988 , 1 ♂, T. Vásárhelyi lgt. (HNHM); ditto, xi.1990 , 2 ♀♀ , G. Csorba lgt. (HNHM); ditto, I.G. 31.668, 25.–28.viii.20l0, day catch, 1 ♀ , 25.–26. viii.2010 , at light, 1 ♀ , J. Constant & P. Limbourg lgt. (ISNB); ditto, 25.–28.viii.2010 , light trap, 1 ♀ , J. Constant & P. Limbourg lgt. (ISNB); ditto, 21°31′N 105°33′E , I.G. 31.933, 25.–30.vii.2011 , 1 ♀ , J. Constant & J. Bresseel lgt. (ISNB); ditto, 21°31′N 105°33′E , 30.vii.2011 , at light, 1 ♀ , J. Constant & J. Bresseel lgt. (ISNB); Tonkin, Guang Yen [= Quang Yen], 26.iii.1915 , 1 ♀ , R.V. de Salvaza lgt. (BMNH). – LAOS : ATTAPEU PROVINCE: Annam Highlands Mts., Dong Ampan NBCA, Nong Fa (crater lake) env., 15°05.9′N 107°25.6′E , ca. 1,160 m a.s.l., 30.iv.–4.v.2010 , 1 ♂ 1 ♀ , J. Hájek lgt. (NMPC). – CHAMPASAK PROVINCE : Bolavens Plateau, ca. 3 km SE Ban Lak 40 (village), coffea plantation / forest, 15°09.8′N 106°08.5′E , 1070 m a.s.l., 9. v.2010 , 2 ♂♂, J. Hájek lgt. (NMPC). – HUA PHAN PROVINCE : Ban Saluei → Phou Pane Mt., 20°12–13.5′N 103°59.5′–104°01′E, 1340–1870 m a.s.l., 15.iv.–15.v.2008 , Lao collectors lgt. (NMPC); Phu Phan Mt., 20°12′N 104°01′E , 1,500–1,900 m a.s.l., 17.v.–3.vi.2007 , 4 ♂♂ 5 ♀♀ , V. Kubáň lgt. (MMBC). – SEKONG PROVINCE : ca. 12 km S Sekong, Tad Faek waterfalls, 15°14.7′N 106°45.l′ E, 118 m a.s.l., at light, 8.–12.v.2010 , 1 ♂, J. Hájek lgt. (NMPC). – VIENTIANE PROVINCE : Lao Pako env., 55 km NE of Vientiane, 200 m a.s.l., 1.–4.v.2005 , 3 ♂♂ 3 ♀♀ , J. Bezděk lgt., Z. Jindra det. (NMPC); Kamphaeng Nakhon Viang Chan prov., 25 km of Vientiane, 1 ♀ , Boudon lgt. (MNHN). – CAMBODIA : KAMPOT PROVINCE: Bokor Nat. Park, Hill Station, scrub, 22.iv.2005 , 1 ♀ , K. Smets & I. Var lgt. (ISNB). – PURSAT PROVINCE : Phnom Samkos W.S., Temple Hill, grassland, bamboo forest, 18.v.2005 , light trap, 1♂, K. Smets & I. Var lgt. (ISNB). – SIEM REAP PROVINCE : Angkor Thorn, 5.iii.2005 , light trap, 1 ♂, D. R. Jump lgt. (ISNB); ditto, viii.2005 , by net, 1 ♂, D. R. Jump lgt. (ISNB). – THAILAND : KAMPAENG PHET PROVINCE: Khiong Lan Nat. Park, 16°07′N 99°16′E , UV pan trap, 6.iv.2003 , 2 ♀♀ , Sites, Vitheepradit, Prommi & Setaphan lgt. (MUUS). – LOEI PROVINCE : Na Haeo, Biol. Station, light trap, 5.–l 2.v.2001 , 4 ♂♂, J. Constant & P. Grootaert lgt. (ISNB); ditto, Na Haeo, river bank, on light trap, l 5.v.2003 , 1♂, P. Grootaert, J. Constant & K. Smets lgt. (ISNB); Na Haeo, forest clearing, on light trap, 16.v.2003 , 1 ♂, J. Constant & K. Smets lgt. (ISNB). – MAE HONG SON PROVINCE : Northern – Pai, 1.–12.v.2001 , 1 ♂ 1 ♀ , R. Kocina lgt., Z. Jindra det. (NMPC); Namtok Mae Surin Nat. Park, Ma Nam Pai, 19°21′N 97°59′E , 310 m a.s.l., 31.iii.2003 , 1 ♂ 1 ♀ , UMC & CMU teams lgt. (MUUS); Pang Mapha, 190°31.159′N 98°14.844′E, 19.iii.2009 , 1 ♀ , R.V. Sites lgt. (MUUS). – NAN PROVINCE : Mae Charin Nat. Park, 18°36′N 100°59′E , 285 m a.s.l., UV pan trap, 22.iv.2003 , 1 ♂ 1 ♀ , Vitheepradit, Prommi & Setaphan lgt. (MUUS). – PHRAE PROVINCE : Wieng Ko Sai Nat. Park, 17°58′N 99°35′E , 350 m a.s.l., pan UV light trap, 29.iii.2003 , 1 ♀ , Sites, Vitheepredit & Prommi lgt. (MUUS). – PITHSANULOG PROVINCE: Phu Hin Rongkhla Nat. Park, on vapor light at guest house, 11.iii.2002 , 3 ♂♂ 4 ♀♀ , Sites, Vitheepredit & Kirawanich lgt. (MUUS); ditto, Phu Hin Rongkhla Nat. Park, Nam Tok Romglao, 16°59′N 101°00′E , 1,190 m a.s.l., 22.iv.2002 , 1 ♂, CMU Team lgt. (MUUS); ditto, Phu Hin Rongkhla Nat. Park, 17°00′N 100°59′E , 1,144 m a.s.l., trap at guest house, 5.v.2003 , 1 ♀ , Vitheepradit, Prommi & Ferro lgt. (MUUS); ditto, Thung Salaeng Luang, 12.v.2004 , 1 ♀ , Vitheepradit & Prommi lgt. (MUUS). – PROCHUAP KHIRI KHAN PROVINCE: Amphur Kui Buri, Forest, Plantation Station, on black light, 12°04′N 99°43′E , 147 m a.s.l., 22.iv.2002 , 1 ♀ , Vitheepradit & Kirawanich lgt. (MUUS). – SURATTHANI PROVINCE: Suratthani env., 1989, 1 ♂, S. Lehman lgt. (ZSMC). – UPPER RATHATARU PROVINCE: Phu Chong Na Yoi Nat. Park, 14°26.l25′N 105°l5.360′E., 1 ♀ , Sites & Vitheepradit lgt. (MUUS); ditto, Phu Chong, Nat. Park, Kaeng Ka Loa, 14°26.178′N 105°16.831′E , 182 m a.s.l., 11.iv.2004 , 1 ♀ , Sites & Vitheepradit lgt. (MUUS). – MALAYSIA : MALAY PENINSULA: FEDERAL TERRITOTY OF KUALA LUMPUR : Kuala Lumpur, at light, 3.xi.1922 , 1 spec. , ex F.M.S. Museum, P. Kment det. (BMNH). – KEDAH : : West Coast, Langkawi Island, 28.iv.1928 , 1 ♂, 29.iv.1928 , 1 ♀ , ex F.M.S. Museum, P. Kment det. (BMNH). – PAHANG : Cameron’s Highlands, 4,000 –5,000 ft [= 1219–1524 m a.s.l.], 10.vi.1935 , H. M. Pendlebury lgt., P. Kment det. (BMNH). – PERAK : Taiping, x.1977 , 3 ♂♂ 7 ♀♀ , P. Pfanner lgt. (MHNG). – SELANGOR : Bukit Kutu, 3,500 ft [= 1067 m a.s.l.], 6.ix.1929 , 1 ♀ , 13.ix.1929 , 1 ♀ , 14.ix.1929 , 1 ♀ , H. M. Pendlebury lgt., P. Kment det. (BMNH). – KALIMANTAN: SABAH : Kinabalu, xi.1978 , 1 ♂, no collector (MNHN) ; Mt. Kinabalu, Nat. Park, 1,500 m a.s.l., 22.–27.iv.1996 , 1 ♀ , S. Snäll lgt. (ZJPC). – SARAWAK : Gunong Mulu Nat. Park, RGS. Mulu Exped., 1 ♀ , J. D. Holloway lgt. (BMNH). – STATE NOT IDENTIFIED: North Borneo, Tawan, on cocoa, 13.ii.1959 , 3 ♂♂, K. L. Leong lgt., M. S. K. Ghauri 1963 det. (BMNH). – SINGAPORE : Singapore, 89-38, no date, 1 spec. , no collecetor (BMNH); Singapore, 96-10, no collector, 1 ♂ 1 ♀ , P. Kment det. (BMNH); Singapore, 97-12, 1 ♀ , H. N. Ridley lgt., P. Kment det. (BMNH); Singapore, 1900-114, H. N. Ridley lgt., P. Kment det. (BMNH). – BRUNEI : Temburong District, ridge NE Kuala Belalong, 125 W m. v. light , ca. 300 m a.s.l., xi.1992 , 1 ♂ 2 ♀♀ , J. H. Martin lgt., P. Kment det. (BMNH); S[ungai] Selanjak, 4463.1432, mangrove, 0 m a.s.l., 6.–7.iii.1984 , 1 ♂, Helps T. P. G. Maj. [lgt.], P. Kment det. (BMNH). – INDONESIA : BALI : Beduqul distr., 1,300 m a.s.l., i.–ii.2005 , 1 ♂♂ 2 ♀♀ , S. Jákl lgt. (ZJPC). – JAVA : JAWA BARAT PROVINCE : Bibidlijan, 10.xi.1937 , 3 ♂♂ 2 ♀♀ , M. E. Walsch lgt. (ZMHB) ; Bogor, no date, 1 ♂, Handschin lgt. (ZMHB); Radjamandala, 1,200 m a.s.l., 6.xi.1937 , 4 ♂♂, M. E. Walsch lgt. (ZMHB); Soekaboemi [= Sukabumi], 1 ♀ , F. Weber lgt. (ZMHB). – JAWA TIMUR PROVINCE : Meru, Betiri Nat. Park, Malangsari villge, 20.v.–5.vi.1996 , 1 ♂, S. Jákl lgt. (ZJPC). – KALIMANTAN : KILIMANTAN SELATAN PROVINCE: S. Kalimantan prov., Loksado 17 km NE, I.G. 31.175, 2 ♀♀ , St. Jákl lgt. (ISNB). – KILIMANTAN TIMUR PROVINCE: Balikpapan, 1947/1948, 1 ♀ , W. A. Mohler lgt. (ZMHB). – MENTAWAI ISLANDS: Nias Island, East coast, Lawlo, 26.ix.1979 , 1 ♂, Ebner lgt. (EHIA). – SUMATRA : RIAU PROVINCE : Indragiri, 1899, 1 ♀ , A. Mechel lgt. (ZMHB) . – SUMATERA BARAT PROVINCE : Bukittinggi, hotel ‘Bukit Tinggi View’, 00°15′30″N 100°21′13″E , 980 m a.s.l., 19.–23.ii.2002 , 1 ♀ , T. Kothe lgt. (ZSMC); Mt. Singgalang, Annai valley env., 600 m a.s.l., v.2006 , 12 ♂♂ 14 ♀♀ , S. Jákl lgt. (ISNB, ZJPC). – SUMATERA UTARA PROVINCE : Dolak Ulu, Kora Kora, 20.ix.1979 , 1 ♂ 1 ♀ , Ebner lgt. (EHIA); Sidikalang, 02°41′51″N 98°18′18″E , 1,250 m a.s.l., 15.–17.ii.2002 , 1 ♂ 1 ♀ , T. Kothe lgt. (ZSMC); near Sipirok, Danau Marsaput, 01°37′50″N 99°20′30″E , 1,435 m a.s.l., 1 ♀ , T. Kothe lgt. (ZSMC) ; S. Pomatang Siantar, 1,100 m a.s.l., 28.–29.x.1992 , 1 ♀ , E. de Bros lgt. (ZMHB). – PHILIPPINES : LUZON: IFUGAO PROVINCE : Mayoyao, viii.1988 , 1 ♂ 1 ♀ , P. Lays lgt. (ISNB). ILICOS NORTE PROVINCE: E of Salsona, 18°05′N 120°54′E , 1,450 m a.s.l., 31.iii.2000 , 2 ♀♀ , L. Dembický lgt. (MMBC) . – MINDANAO: SOUTH COTOBATO PROVINCE: Lake Sebu, Mt.Tasaday, 6°18′N 124°42′E , v.–vi.1994 , 1 ♂, P. Lays lgt. (ISNB) ; Lake Sebu, 6°13′N 124°42′E , 700 m a.s.l., at light trap (neon), 1.–15.i.1998 , 1 ♂ 2 ♀♀ , P. Lays lgt. (ISNB) . LANAO DEL NORTE PROVINCE : Iligan, 1 ♀ , W. Schulze lgt. (ZSMC) . – BUKIDNON PROVINCE : Bukidnon, 1.–10.x.1988 , 3 ♂♂ 1 ♀ , K. Černý lgt. (MMBC). – NEGROS ISLAND: NEGROS ORIENTAL PROVINCE: Camp Lookout, Dumaguete, 15.ii.–15.iv.1961 , 1 ♂, T. Schneiria & A. Reyes lgt. (AMNH). – PALAWAN : Irawan Forest, 1000 ft , swept, B.M. 1988-2, i.–ii. 1988 , 1 ♀ , J. H. Marting lgt. (BMNH); Mantalingajan Range, Pinigisan, approx. 600 m a.s.l., 31.viii.1961 , 1 ♂, 8.ix.1961 , 1 ♂, 9.ix.1961 , 2 ♂♂ 2 ♀♀ , all Noona Dan Expedition (ZMUC); Pa1awan, Irawan Forest, swept, 1,000 ft [= 305 m a.s.l.], i.–ii.1988 , 1 ♀ , J. H. Marhn lgt. (BMNH). – TAWI TAWI ISLANDS: Tarawakan, north of Batu Batu, 14.xi.1961 , 3 ♀♀ , Noona Dan Expedition (ZMUC); ditto, Lapid Lapid at Manalik Channel, 19.xi.1961 , 1 ♀ , Noona Dan Expedition (ZMUC). Distribution. Japan (Honshu, Kyushu, Ryukyus) ( SCOTT 1874 , DISTANT 1883 , UHLER 1897 , BLÖTE 1931 , MANNA et al. 1985 , KERZHNER 2001 , KOHNO et al. 2002 , TOMOKUNI 2006 ; this paper), Korea (South, Chejudo Island) ( LEE & KWON 1991 , KWON et al. 2001 , KERZHNER 2001 ), China ( Fujian , Guangdong , Guizhou , Hubei , Hunan , Jiangsu , Jiangxi , Sichuan , Yunnan , Xizang , Zhejiang , Hainan ) ( STÅL 1870 , WALKER 1873 , DISTANT 1903a , LIU 1981 , HUA 2000 , KERZHNER 2001 ; this paper), Taiwan ( ESAKI 1926 , BLÖTE 1931 , KERZHNER 2001 , RÉDEI et al. 2009 ; this paper), Afghanistan ( KERZHNER 2001 , no details; this paper), Pakistan ( Khyber Pakhtunkhwa , Punjab ) ( AHMAD & ABBAS 1987 ; this paper), India ( Andhra Pradesh , Arunachal Pradesh , Assam , Bihar , Haryana , Himachal Pradesh , Jammu and Kashmir , Karnataka , Madhya Pradesh , Meghalaya , Tamil Nadu , Uttarakhand , West Bengal ) ( DISTANT 1879a , 1903a ; MAXWELL- LEFROY 1909; SCHMIDT 1931 ; MANNA et al. 1985 ; ZAMAL & CHOPRA 1990 ; CHAKRABARTY et al. 1994 ; SAHA & BAL 2007 ; STEHLÍK 2007a ; CHANDRA et al. 2012 ; CHANDRA & KUSHWAHA 2013 ; SINGH & BANYAL 2013 ; this paper), Bhutan (new record), Nepal ( STEHLÍK 2004 ; this paper), Sri Lanka ( WALKER 1873 , KIRBY 1891 ; this paper), Bangladesh ( WALKER 1873 ), Myanmar ( DISTANT 1879b , 1903a ; this paper), Vietnam (new record), Laos ( STEHLÍK 2005a ; this paper), Cambodia (new record), Thailand ( DISTANT 1903b , STEHLÍK & JINDRA 2003 ; this paper), Malaysia (Peninsular Malaysia: Federal Territory of Kuala Lumpur , Kedah , Pahang , Penang , Selangor ; Kalimantan: Sabah , Sarawak ) ( WALKER 1873 , BREDDIN 1900 ; DISTANT 1903b ; this paper), Singapore (new record), Brunei (new record), Indonesia ( Bali , Java , Kalimantan , Mentawai Islands: Nias, Sumatra ) ( BURMEISTER 1834 , 1835 ; STÅL 1855 , as Ph. bimaculata , 1870 ; WALKER 1873 ; LETHIERRY 1888 ; BREDDIN 1901b ; DISTANT 1903a ; BLÖTE 1931 ; SCHMIDT 1931 ), Philippines ( Basilan , Luzon, Mindanao, Negros, Palawan , Tawi Tawi Island) ( BURMEISTER 1834 , STÅL 1870 , WALKER 1873 , TAEUBER 1927 ; this paper). Distributional records from Australia and Australian Region by BREDDIN (1900 , 1901a ), CASSIS & GROSS (2002) , and VOIGT (2006) belong to Ph. gutta famelica (see e.g., STEHLÍK 2007a ). The records of Ph. gutta from Sulawesi ( BREDDIN 1901a ), Timor ( Blöte 1931 ), and Halmahera ( BREDDIN 1900 ) apparently belong to Ph. gutta famelica as well. The record of Ph. gutta famelica from China : Sichuan by VOIGT (2006) represents a misidentification of Ph. gutta gutta .