Rediscovery and redescription of the centipede Paracryptops inexpectus Chamberlin, 1914, with an account of the genus (Scolopendromorpha: Cryptopidae: Cryptopinae)
Author
Junior, Amazonas Chagas
Author
Shelley, Rowland M.
text
Zootaxa
2004
475
1
8
journal article
10.5281/zenodo.157786
c1cd13ed-07de-4ea3-9b6a-74522261bd3b
11755326
157786
Genus
Paracryptops
Pocock, 1891
Paracryptops
Pocock, 1891
:227
; 1894:316.
Kraepelin, 1903
:59
–60.
Verhoeff, 1907
:250
.
Chamberlin, 1914
:158
.
Attems, 1930
:244
. B cherl, 1939:240; 1941:74; 1974:123.
Khanna, 1994
:460
; 2001:211.
Schileyko, 2002
:482
.
Type
species
.
P. weberi
Pocock, 1891
, by monotypy.
Diagnosis
. Anterior margin of coxosternum with two rounded, segregated lobes; claws of prehensors short, barely extending beyond levels of inner margins of trochanteroprefemora, well separated from opposite member in resting position.
Components
. Five species have been described, some of which may be synonymous:
P. w e b e r i
,
P. breviunguis
,
P. inexpectus
,
P. i n d i c u s
, and
P. spinosus
.
Distribution
. Southern and southeast Asia (
India
,
Malaysia
[Sabah vicinity],
Vietnam
,
Singapore
,
Indonesia
[Java,
Flores
, and Sumba], and
Papua New Guinea
); South
America
(
Guyana
); Lesser Antilles (
Dominica
) (
Fig. 1
).
Schileyko (2002)
also cited the genus from
China
, but a thorough search of the second author’s library did not produce a documentation from
China
, and we do not know the source of this citation.
China
should be deleted from the generic range until confirmed with a definite record.
Remarks
.
Paracryptops
is clearly a valid genus in the Cryptopinae; the lobed anterior margin of the coxosternum and the short prehensor claws are reliable generic characters (
Pocock, 1891
,
1894
;
Attems, 1930
;
Jangi & Dass, 1978
). Only around
20 specimens
total have been collected, and all the component species occur in southern/southeast Asia and the East Indies except for the isolated occurrence of
P. inexpectus
in the West Indies and northern South
America
, some
10,475 mi
(
16,752 km
) to the east (
Fig. 1
). This distribution pattern suggests that
P. inexpectus
is an allochthonous species of anthropogenic origin rather than an indigenous New World taxon.
As
the largest community and only true city on
Dominica
, Roseau probably harbors a number of introduced centipede and millipede species, which typically predominate in urban environments, suggesting that the specimens of
P. inexpectus
are exogenous, although the source area is unknown. Likewise, the two localities of
P. spinosus
, Delhi and New Delhi, are large cities that surely harbor many introduced species, and the intercepted specimen of
P. w e b e r i
came from such a place,
Singapore
; indeed five of the 16 samples of
Paracryptops
(31% of the total) came from urban environments, so the genus is intimately associated with man. Additionally, representatives of
Paracryptops
have twice been intercepted in quarantines, in Honolulu (from
Singapore
) and Washington, DC (from
Guyana
), and the
type
of
P. spinosus
was found under pots in a plant nursery, where it was likely carried by man. The distribution pattern of four species in southern/southeast Asia and one thousands of miles away in the eastern Caribbean and northeastern South
America
defies geographic logic unless the latter is introduced from the former area. This possibility is enhanced by the small size and cryptic nature of the centipedes, and the fact that, according to the descriptions, there are few differences between the species. We suspect that
Paracryptops
actually consists of less than five species, perhaps only one or two, that have been unknowingly introduced by man to other locations and then discovered and redescribed as new species. A full generic revision, including examinations and comparisons of all specimens, especially the
types
, is needed and would provide clues to the origins of the species, but this is beyond the scope of the present study. We therefore retain all five species and redescribe
P. inexpectus
based on the
holotype
and the two Dominican individuals.