Three cryptic Anaplecta (Blattodea, Blattoidea, Anaplectidae) species revealed by female genitalia, plus seven new species from China
Author
Zhu, Jing
College of Plant Protection, Southwest University, Beibei, Chongqing 400715, China
Author
Zhang, Jiawei
College of Plant Protection, Southwest University, Beibei, Chongqing 400715, China
Author
Luo, Xinxing
College of Plant Protection, Southwest University, Beibei, Chongqing 400715, China
Author
Wang, Zongqing
College of Plant Protection, Southwest University, Beibei, Chongqing 400715, China
Author
Che, Yanli
College of Plant Protection, Southwest University, Beibei, Chongqing 400715, China
shirleyche2000@126.com
text
ZooKeys
2022
2022-01-04
1080
53
97
http://dx.doi.org/10.3897/zookeys.1080.74286
journal article
http://dx.doi.org/10.3897/zookeys.1080.74286
1313-2970-1080-53
86DAAF2DC098452BB3EA51D84EB5855E
B74322796C11528189E0ABBEE84DD187
Anaplecta condensa Zhu & Che
sp. nov.
Figures 2C, D
, 12
, 16G-I
Type material.
Holotype
: China • male; Guizhou Prov., Libo County, Jiaou Village;
25°30.06'N
, 107°67.02'E; 11 June 2019; Lu Qiu, Wen-Bo, Deng, leg.; SWU-B-B-A060121.
Paratypes
: China • 3 males and 1 female; same data as holotype; SWU-B-B-A060122 to 060125 • 2 males; Guangxi Prov., Guiping City; 31 May-2 June 2014; Shun-Hua Gui, Xin-Ran Li, Jian-Yue Qiu, leg.; SWU-B-B-A060126 and 060127.
Diagnosis.
This species is very similar to
A. omei
, but can be distinguished as follows: 1) paraprocts both extending backward in
A. condensa
sp. nov., while only the left extending backward in
A. omei
; 2) R1 needle-shaped in
A. condensa
sp. nov., while bifurcated in
A. omei
; and 3) intercalary sclerite of
A. condensa
sp. nov. very small, filamentous, while that of
A. omei
is spindle-shaped.
Etymology.
The specific epithet is derived from the Latin word
condensus
, referring to the paraprocts with dense spines on curly posterior margin.
Measurements (mm).
Male: pronotum length
x
width: 1.36-1.39
x
1.78-1.84, tegmina length: 4.93-5.39, overall length: 5.92-6.59. Female: pronotum length
x
width: 1.29
x
1.73, tegmina length: 4.75, overall length: 5.82
Description.
Coloration
.
Body brown (some individuals from Guiping yellowish brown) (Fig.
2C, D
), face dark brown (Fig.
12A, B
). Antennae and maxillary palpus brown (Fig.
12D
). Pronotum dark brown, lateral edges nearly hyaline (Fig.
12C
). Tegmina yellowish brown, anal field and base of mediocubital field slightly darker (Fig.
12E
). Hind wings infuscate, costal field and appendicular field darker than remaining parts (Fig.
12F
). Center of abdominal sterna yellow, gradually darkening to dark brown to edges, legs, and cerci dark yellowish brown (Fig.
12B
).
Figure 12.
Anaplecta condensa
Zhu & Che, sp. nov. holotype (GZ4), male SWU-B-B-A060121
A
habitus, dorsal view
B
habitus, ventral view
C
pronotum, dorsal view
D
head, ventral view
E
tegmina
F
wings
G
supra-anal plate, ventral view
H
subgenital plate, dorsal view
I
left phallomere, dorsal view
J, K
right phallomere
J
dorsal view
K
ventral view. Scale bars: 2 mm (
A-F
); 0.5 mm (
G-K)
. Abbreviations:
afd
anal fold,
A[1
] the anterior one of the anal vein,
A[s
] the other element of vannal vein,
cfd
cubitus fold,
CuA
cubitus anterior,
CuP
cubitus posterior,
L1, L2, L3
sclerites of the left phallomere,
L2v
L2 ventral,
L2vm
median sclerite,
M
media,
Pcu
postcubitus,
R
radius,
RA
radius anterior,
RP
radius posterior,
R1, R2, R3
sclerites of the right phallomere,
ScP
subcostal posterior.
Head and thorax
.
The distance between antennal sockets slightly narrower than interocular space. Fifth maxillary palpus nearly oval, slightly thicker and wider than others (Fig.
12D
). Pronotum semicircular, anterior margin arched, the center of posterior margin protrudes slightly (Fig.
12C
). Tegmina with indistinct veins, radius posterior veins of hind wings slightly indistinct, without transverse veins between M and CuA (Fig.
12E, F
). Front femur Type B2 (Fig.
12B
). Pulvilli absent, tarsal claws symmetrical.
Male genitalia
.
Paraprocts both extend backwards and with dense spines on curly posterior margin (Fig.
12G
). Subgenital plate sub-rectangular, the center of anterior and interstylar margins nearly straight. Styli long, so is the distance between them (Fig.
12H
). L1 subcircular, with curved and long filamentary structure. L2v curved, bifurcated, with a sharp horn. L2vm broad. L3 extremely robust, uncinate part blunt (Fig.
12I
). R1 needle-shaped, the proximal part slightly curved (Fig.
12J
). R2 irregular, weakly sclerotized. R3 slightly curved, sheet-like (Fig.
12K
).
Female genitalia
.
Supra-anal plate nearly symmetrical, very blunt and round. Paraprocts broad, hind margin blunt, not extending to the posterior margin of supra-anal plate. Intercalary sclerite small, nearly filamentous. First valve robust. Second valve small, basally fused. Third valve broad. The anterior margin of anterior arch slightly curled, with a hook-shaped protrusion (Fig.
16G-H
). Basivalvula broad, with dense punctuations, except for left lateral and anterior margin (Fig.
16G
). Vestibular sclerite broad, slightly curled, sheet-like. Laterosternal shelf slightly sclerotized, lateral margin straight (Fig.
16I
).
Distribution.
China (Guizhou, Guangxi).