Onthophagus (Palaeonthophagus) medius (Kugelann, 1792) — a good western palaearctic species in the Onthophagus vacca complex (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini)
Author
Rössner, Eckehard
Author
Schönfeld, Joachim
Author
Ahrens, Dirk
text
Zootaxa
2010
2629
1
28
journal article
10.5281/zenodo.198282
415ebeff-395d-4ba4-934e-697e64bb2664
1175-5326
198282
Onthophagus
(
Palaeonthophagus
)
medius
(
Kugelann, 1792
)
(revised status)
Copris medius
Kugelann, 1792
: 271
(
Type
locality: "Preußen").
Scarabaeus medius
Panzer, 1796
: 1
(
Type
locality:
Germany
).
Copris affinis
Sturm, 1800
: 102
(
Type
locality: "Michaelsberg bei Herspruck" [probably Bavaria]).
Onthophagus vacca
var.
vicinus
Mulsant, 1842
: 133
(
Type
locality:
France
).
Onthophagus vacca
var.
intermedius
Mulsant, 1842
: 134
(
Type
locality:
France
).
Onthophagus vacca
var.
propinquus
Mulsant, 1842
: 134
(
Type
locality:
France
).
Onthophagus vacca
var.
sublineolatus
Mulsant, 1842
: 134
(
Type
locality:
France
).
Onthophagus vacca
var.
basalis
Mulsant, 1842
: 134
(
Type
locality:
France
).
Onthophagus confluens
Gistel, 1857
: 61
(
Type
locality:
Germany
).
Type
material examined:
Neotype
(
Copris medius
,
here designated
) [Figs. 2L, 2P, 2AF, 2AJ, 3A, 3B, 3AG, 3AP]: "Lyck, Ostpreussen [black handwritten] B.v.Bodemeyer [black printed on white paper] |
Onthophagus vacca v. medius
[black handwritten on white paper] |
Neotypus
Copris media
Kugelann, 1792
, E. Rößner, 2008 [black handwritten on red paper] |
Onthophagus (Palaeonthophagus) medius (
Kugelann, 1792
)
, det. E. Rößner, 2008 [black handwritten on white paper]" (ɗ,
ZMHB
) [specimen glued on carton, genital extracted].
Denmark
: "Helnaes, Bobakker
8.v.1999
| DK.F.NG61, P. Jörum leg." (2 ΨΨ, CPJ).- "Rösnaes (Rösnaesgd.)
11.vi.2005
| DK.NWZ.PG17, P. Jörum leg." (1 ɗ, 1 Ψ, CPJ).- "Mögelskår
19.v.2004
| DK.EJ.NH90, P. Jörum leg." (2 ΨΨ, CPJ).- "Kopenhagen | coll. Leonhard" (2 ɗɗ, 2 ΨΨ,
DEI
).
Netherlands
:
"Wateringen,
4.V.1954
, R. Bouwer" (1 ɗ, CJS).- "Opheusten,
20.VI.1957
, R. Bouwer" (1 Ψ, CJS).- "Maastricht, 5.1921, [leg.] Rüschkamp, Sammlung F. Rüschkamp. Eing. Nr. 1.1964" (2 ΨΨ,
ZMFK
).
Luxemburg
:
"
Luxemburg
, Differdingen, 4.[19]15, P. Heymes" (1 Ψ, coll. Heymes,
NME
).- "
Luxembourg
: Remich, Erpel D’Ange, Chatea Dames
29.viii.1998
, ex horse dung, coll. M.V.L. Barclay" (4 ɗɗ, 11 ΨΨ,
NHML
).- "Grevenmacher, Manternach, 0 2.06.2001, leg. Dirk Rohwedder" (1 ɗ, CDR).- "Grevenmacher, East-Gonderange, Schafkot, 0 3.06.2001, leg. Dirk Rohwedder" (1 Ψ, CDR).- "Gonderange, 0 1.06.2001, leg. K. Hadulla" (1 Ψ, CKHa).
Belgium
:
"
Belgium
" (1 ɗ, 1 Ψ, coll. Fry,
NHML
).- "Antwerpen
29.5.1898
." (1 ɗ,
NMP
).
Germany
:
"Mecklenbg.: Klein Schmölen, Pferdeweide an der Löcknitz,
2833/IV
,
01.V.2005
, Pferdekot, leg. E. Rößner" (5 ɗɗ, 5 ΨΨ, CER).- dto., nur: "
25.IV.2006
" (4 ɗɗ, CER).- "Lebus/
O
. 23.5.[19]72" (1 ɗ, 1 Ψ,
ZMHB
).- "Lebus, Oderbruch,
DDR
,
21.V.1982
, leg. U. Heinig" (1 Ψ, CER).- "Oderhänge bei Lebus,
DDR
,
28.V.1988
, leg. Schröder" (1 ɗ, 1 Ψ, CER).- "Teichhorn, Volksen [bei Einbeck], 5.6.[19]55, Kuhmist, Priefert & Helmke lg." (1 ɗ,
ZMHB
).- "Bottendorf [wahrscheinlich: bei Obernholz], 23.6.[19]55, Lohse" (1 Ψ,
ZMHB
).- "Baden b. Bremen 13.5.[19]51, 53, Kerstens" (1 ɗ, CAB).- "Bremen, Baden
11/6 1950
, J. D. Schroeder" (1 ɗ, CAB).- "Badener Berge.
8.IV.
[18]94. Achim. Hannover, coll. Künnemann" (1 ɗ,
DEI
).- dto., nur "
24.IV.98
" (1 Ψ,
DEI
).- "Lindhorst, Kr. Stadth. Hannover,
IV.1951
, leg. E. Gersdorf" (1 Ψ, CJS).- "Geestgottberg, ST Altmark, Elbedeich b. Brücke, Schafkot
17.05.2006
[leg.] Malchau" (1 ɗ, 2 ΨΨ, CWM).- "
Germania
sept., Gatersleben (Quedlinburg),
22.6.1952
. 127. [leg.] Dorn" (3 ɗɗ, 3 ΨΨ,
ZMHB
).- "Dornburg [bei Zerbst], JL, ST, Gommern, Weide östl. Ortsrand,
24.05.2007
, Malchau" (1 ɗ, 3 ΨΨ, CWM).- "Neugattersleben, ST, BBG, Bodegrünland bei A 14, Pferd Schaf,
16.05.2007
, Malchau" (1 ɗ, 2 ΨΨ, CWM).- "Breitenhagen: Sommerdeiche,
18.IV.2005
, leg. E. Rößner" (2 ɗɗ, 2 ΨΨ, CER).- "Sachsen-Anhalt: NSG Lämmerberg und Vockenwinkel bei Wormsleben, 4435/IV,
29.V.1998
, leg. E. Rößner" (2 ɗɗ, 4 ΨΨ, coll. CER; 3 ɗɗ, CHKa).- "Sachsen-Anhalt: Wormsleben, 4435/IV,
26.V.1999
" (2 ɗɗ, 1 Ψ, CER).- "Sachsen-Anhalt, Dabrun bei Wittenberg, 4142/3, Rinderkot,
18.05.2002
Bäse leg." (1 ɗ, 1 Ψ,
CWB
).- "Sachsen-Anhalt 4244/2, Premsendorf bei Jessen, an Pferdekot,
16.05.2005
K. Bäse leg." (1 ɗ, 1 Ψ,
CWB
).- "Beesenlaublingen, ST, BBG, A 14 Brücke, Saaledeich, 0 3.05.2007, Malchau" (1 Ψ, CWM).- "Thuringia MTB 4531/4, Numburg- West, Kelbra,
20.V.
[20]05, Salzwiese, Sw B, leg. H. Sparmberg" (1 Ψ, CHS).- "Thuringia MTB 4531/4 Kelbra, Numburg, West,
27.IV.2007
Hf leg. H. Sparmberg" (2 ΨΨ, CHS).- "Thuringia,
17.V.2002
, 4832/III, Haßleben, Rinderweide, trocken, Fangart: Hf, leg. H. Sparmberg" (1 Ψ, CHS).- "Kyffhäuser Schlachteberg
21.04.1989
, R. Peschel" (1 Ψ,
CRP
).- "Bad Frankenhausen, A.06.1983, leg. E. Rößner" (1 Ψ, CER).- "
BRD
/ Thür./ Kyffhäuser, Udersleben 250 NN, U. Hämling,
28.4.1994
, leg. U. Schaffrath" (1 ɗ, 1 Ψ,
CUS
).- "Thuringia MTB 4632/IV, Esperstedter Ried, Hf,
17.VI.
[20]05, Gebiet A, leg. H. Sparmberg" (1 ɗ, 1 Ψ, CHS).- "Rastenberg: Kapellenberg, 4834/II, HF,
08.VI.2008
, leg. H. Sparmberg" (3 ɗɗ, CHS).- "Nöda: Rinderweide, Feuchtwiese, 4932/I, HF,
14.VI.2008
, leg. H. Sparmberg" (1 ɗ, CHS).- "Meining[en]. (Heyd.)" (2 ɗɗ, 1 Ψ, coll. Heyden,
DEI
).- "Saxonia Provincia, Eilenburg, Hainichen Nord, 13.4.1953.32, [leg.] Dorn" (2 ɗɗ, 2 ΨΨ,
ZMHB
).- "14.5.[19]56, Naunhof [Großenhain], [leg.] Hieke" (1 ɗ,
ZMHB
).- "Meißen, Zadel, 24.6.[19]70" (1 Ψ,
ZMHB
).- "D: Saxonia cent., Köllitzsch/Elbe
29.V.2001
Grünland, Projekt Uni Jena BF" (1 ɗ, 1 Ψ,
NME
).- "Düben Kirsch" (1 Ψ,
SMTD
).- "Wehlen Märkel" (1 Ψ,
SMTD
).- "Grunaer Wald, Düb. Heide 3.5.[19]53, L. Dieckmann, Pferdemist" (2 ɗɗ, 2 ΨΨ,
DEI
).- "D, Umg. Schlüchtern, Elm, in Kuhdung, 500 NN,
11.6.1989
, leg. Schaffrath" (1 Ψ,
CUS
).- "Lampertheim/ Rhein, S-Hessen D.,
1.5.1999
, leg. G. Müller" (1 Ψ,
CUS
).- "Darmstadt 29.4.-
30.7.1949
E. Möhn leg." (1 Ψ,
SMNS
).- "Hofgeismar, Hess.-Nass.
4.V.
[19]30" (3 ɗɗ, 1 Ψ,
SMNS
).- "Wiesbaden. | Collektion Kirschbaum" (1 Ψ,
NSMW
).- "Frankft. Hyd." (4 ɗɗ, 2 ΨΨ, coll. Heyden,
DEI
).- "Rheinprovinz, Rees, 11.-13.6.[19]35, K. Ermisch." (2 ɗɗ,
ZMHB
).- "D., R.-Pf., UG Mainz, Nackenheim, in Pferdeäpfeln, 100 NN, 22.-
24.4.1990
, leg. U. Schaffrath" (1 Ψ,
CUS
).- "D. Rheintal, Bacharach, Steeg Weinberge, F. Köhler
22.05.1996
" (1 ɗ,
ZMHB
).- "D,
RP
, Nahe-Glan-Gebiet, Steckweiler, Solzenberg
280 m
24.04.2001
, leg. F. Köhler" (5 ɗɗ, 2 ΨΨ, CHB).- "Rheinland-Pfalz: Steckweiler (im Alsenztal), NSG Stolzenberg,
24.IV.2001
, leg. Frank Köhler" (26 ɗɗ, 10 ΨΨ, CER).- "D – RP-LS, Mosel, Kobern, Bellbach-Tal
100 m
, 0 9.05.2002, leg. J. Beller" (1 Ψ, CHB).- "Ger.,
28.V.1978
, Rh.-Pf./ Mosel, Trier, Petrisberg. Scheuern leg." (1 ɗ, CJS).- "D – Rheinl.-Pfalz/ Südpfalz, Schweighofen (SÜW) beim Bahnhof,
24.V.2006
, leg. L. Schmidt" (2 ɗɗ, 4 ΨΨ, CLS).- "D/ BW/ Breisach Oberbergen, Schelinger Weide, Kaiserstuhl,
4.6.2000
, leg.: Willers" (1 ɗ,
ZMHB
).- "Kaiserstuhl, Schelingen, 0 4.06.2000, K. Renner" (1 ɗ, CKR).- "Rheinland/Pfalz, Schweighofen, Schafdung in Falle,
19.05.2007
, leg. Dirk Rohwedder" (2 ɗɗ, CDR).- "Düsseldorf, L,
5.6.1962
" (1ɗ, coll. Koch,
ZMFK
).- "Düsseldorf, Z,
17.6.1964
" (1 ɗ, coll. Koch,
ZMFK
).- "Worring[er]. Bruch, [leg.] H. Appel
18.5.1957
, ex Coll. Appel" (1 ɗ, 1 Ψ, coll.
CAG
Wuppertal,
ZMFK
).- "Rh. Ufer b. Köln, [leg.] Appel
2.5.1965
, ex Coll. Appel" (1 Ψ, coll.
CAG
Wuppertal,
ZMFK
).- "Bienwald, Kuhweide
500 m
N Kapweyer,
49°2’4" N
8°1’20" E
,
24.05.2008
, leg. J. Schönfeld" (1 ɗ, 1 Ψ, CJS).- "Bienwald, Pferdeweide
1 km
E Büschelberg, W der B 9,
49°1’17" N
8°11’7" E
,
22.05.2008
, leg. J. Schönfeld" (1 Ψ, CJS).- "Bienwald, Kuhweide
500 m
W Neulauterburg, nahe der Lauter,
48°58’42" N
8°10’8" E
,
23.05.2008
, leg. J. Schönfeld" (1 Ψ, CJS).- "Kaiserswerth, Rheinufer (südl.),
14.5.1919
, Kirsch l." (4 ɗɗ, coll.
ARK
,
ZMFK
).- "Düsseldorf,
29.5.1928
, [leg.] C. Henseler" (1 ɗ, coll.
ARK
,
ZMFK
).- "Elberfeld, [leg.] Geilenk[euser]." (3 ɗɗ, coll.
ARK
,
ZMFK
).- "Niederrhein, Rees, [leg.] Horion, K., 6.1935" (1 ɗ, 1 Ψ, coll.
ARK
,
ZMFK
).- "Rees, Nrh.,
Juni 1935
, [leg.] P. Müller, Sammlung M. Schmaus, Eing.Nr. 2, 1979" (1 ɗ, coll.
ARK
,
ZMFK
).- "Acker, Bonn,
9.5.1927
, [leg.] Rüschkamp" (1 Ψ, coll.
ARK
,
ZMFK
).- "Stobeln,
7.5.1928
, [leg.] C. Henseler" (1 Ψ, coll.
ARK
,
ZMFK
).- "Kastellaun,
28.6.1949
, [leg.] M. Schmaus" (1 Ψ, coll.
ARK
,
ZMFK
).- "Hessen, Wetterau, 4.1935, [leg.] Schoop 09" (1 ɗ,
ZMFK
).- "Schelingen/Kaiserstuhl, Rheinaue, 28.4.[19]96 leg. Esser" (1 Ψ, CAB).- "Bavaria, Erlangen" (2 ɗɗ, 3 ΨΨ,
ZMHB
).- "Ingolst[adt]. 22.5.[19]02" (1 Ψ,
SMTD
).- "München, Coll. Kraatz" (1 ɗ, 1 Ψ,
DEI
).- "Monachium [= München],
var. medius Panz.
" (1 ɗ,
ZSM
).- "Monachium,
var. affinis Sturm.
" (1 ɗ,
ZSM
).- "Straubing 8.[19]11" (1 ɗ,
ZSM
).- "Geisenfeld
21.V.
[19]36 | Sammlung
O
. Bühlmann" (1 ɗ,
ZSM
).- "Schleissheim 7.6.[19]04 | München H. Kulzer" (1 ɗ, 1 Ψ,
ZSM
).- "Hartmannshf. [= Hartmannshof], 19.6.[19]06 | München H. Kulzer" (1 ɗ, 1 Ψ,
ZSM
).- "München Umg., Obb." (1 ɗ,
ZSM
).
Poland
:
"Posen [= Poznań], [leg.] Paulisch" (1ɗ,
ZMHB
).- "Ohlau [= Oława] | Coll. Letzner" (1 ɗ,
DEI
).- "Ost-Pr[eußen]." (1 Ψ,
ZMHB
).- "Silesia Kirsch" (1 ɗ,
SMTD
).
Austria
:
"Neusiedlersee,
16.6.1936
, Burgenland, J. Wirthumer" (1 ɗ, 1 Ψ,
NMP
).- "
Österreich
Wien | ex coll. Dr. Kallert Hamburg" (2 ɗɗ,
NMP
).- "Präwald [Oberösterreich, südwestlich Windischgarsten], Kr, 26.6.[19]08" (1 Ψ,
ZMHB
).- "Burgenland, Neusiedl" (1 ɗ, 2 ΨΨ,
ZMHB
).- "Neusiedler See,
25.5.1975
" (1 ɗ,
ZMFK
).- "Umg. Wien, [leg.] Ad. Hoffmann, ex Coll. Missionshaus SVD Steyl, Eing. Nr. 3/59" (4 ΨΨ,
ZMFK
).- "Neusiedler See u. Umg. A 6 [19]63 leg. Witzgall" (1 Ψ,
CRP
).- "Neusiedl. See u. Ug.,
Austria
, Dr. Lechner" (1 ɗ, CAB).- "12.7.[19]73 Burgenland, Parndorf/Hutweide" (1 Ψ, CAB).- "A. NSS Apetlon
6.5.1986
leg. Hirgstetter" (1 Ψ,
CRP
).- "4.4.[19]74 Nähe Apetlon,
Austria
, Neusiedler See, Fery leg." (2 ɗɗ, 1 Ψ, CHF).- "15.5.[19]76, Pamhagen (
Austria
), Neusiedler See, Pferdekot, Fery leg." (1 ɗ, CHF).- "
Hungaria
, Leitha-Geb., F. Blühweiss" (2 ΨΨ,
ZMHB
).- "Illmitz Brgld. 11.5.[19]66 Folwaczny" (5 ɗɗ, 1 Ψ,
SMNS
).- "Neusdl. See
25.5.1975
" (2 ΨΨ,
SMNS
).- "Aust. | 26960" (4 ɗɗ, 2 ΨΨ,
ZMHB
).- "Austria-Burgld., leg. Hebauer | Neusiedl. See,
25.5.1975
" (3 ΨΨ, CJS).- "A.i. Leitha-Geb., Ad. Hoffmann" (1 Ψ,
SMTD
).- "
2.VI.1928
. Umgebung Apetlon am Neusiedler See" (1 Ψ,
SMTD
).- "Neusiedler See, leg. R. Zimmermann 1941, Schmidt" (3 ɗɗ, 2 ΨΨ,
SMTD
).- dto., nur "
Juli 49
" (1 Ψ,
SMTD
).- "Bündten, coll. Stierlin" (1 ɗ,
DEI
).- "Villach" (1 ɗ,
ZSM
).- "Neusiedler See, leg. Zimmermann 1941, [coll. H.] Schmidt" (1 Ψ,
ZSM
).- "
Austria
Bgld. Parndorf, Hüdepohl 5.[19]69" (1 Ψ,
ZSM
).- "Neusiedler See,
V.1953
, [leg.] Dr. H. Lechner" (1 ɗ, CLS).
Czech Republic
:
"Boh., Louny/Oblik 2.5.[19]62 A. Hoffner leg." (1 ɗ, 1 Ψ,
NMP
).- "Boh., Louny env.
30.IV.
[19]
61 V.
Balthasar leg." (4 ɗɗ, 5 ΨΨ,
NMP
).- "Závist, 21… Zeman" (1 ɗ,
NMP
).- "Env. de Praque,
Bohême
centr., Coll. Obenberger" (1 ɗ,
NMP
).- "Radotin
21.5.1927
" (1 ɗ,
NMP
).- "
Bohemia
, Jos. Cêrný | Pardubice
18.5.1909
| Mus. Pragensde, coll. Kracik” (1 Ψ,
NMP
).- "Moravia-1934, Veseli n[ad]. Mor[avon]. VI. [leg.]
O
. Kodym" (1 Ψ,
NMP
).- "Prossnitz [= Prostĕjov], [leg.] K. Kyselý” (1 ɗ,
NMP
).
Slovakia
:
"Bratislava, Petržalka, [leg.] Roland" (1 Ψ,
NMP
).- "Slov. 3-4.[19]39, Bratislava" (1 Ψ,
NMP
).- "Slov. mer., Stúrovo, 28.-
30.7.1955
, V. Balthasar leg.” (1 ɗ,
NMP
).- "
Slovakia
merid., Štúrovo | Parkán | [leg.]
O
. Kavan, 24.4.[19]49" (2 ɗɗ, 4 ΨΨ,
NMP
).- "
Slovakia
1935. Plavec[ký] Sv[ätý] Mikuláš
27.V.
, [leg.]
O
. Kodym" (1 ɗ,
NMP
).- dto., nur "
5.VI.
" (1 ɗ,
NMP
).- "
Slovakia
merid., Kováčovské vrchy,
O
. Kavan| Kováčar, 27.4.[19]49" (1 ɗ, 5 ΨΨ,
NMP
).- "
Slovakia
merid., Kováčovské vrchy,
O
. Kavan | Helemba, 22.4.[19]49" (1 ɗ,
NMP
).- "Slovakei, Pannonisches Becken, Zádiel Umg.
200 m
, 0 1.07.1970 leg. J. Hadiel" (1 ɗ, CHB).- "Zadiel
1.VII.
[19]70" (1 ɗ,
SMNS
).- "
Slovakia
, Štúrovo-Bet.k., 29.4.[19]54, Kr. Pospišil leg." (1 ɗ,
SMTD
).- "
Slovaquie
, Košice V.[19]37 coll. Pospišil" (1 Ψ,
SMTD
).- "
Slowakei
, Silicka planina 12.-
22.7.1993
, leg. Richter" (1 Ψ, CWR).
Hungary
:
"Hungarie c. or., Pécz, J. Pradáč leg." (1 Ψ,
NMP
).- "Hungarie centr., Békéscsaba env.,
V.1947
, lgt. Škpt Havelka” (1 Ψ,
NMP
).- "Ujpest,
Ungarn
| Mus. Pragense Coll. Kracik" (1 ɗ, 1 Ψ,
NMP
).- "
Hungaria
, Esztergom, E. Bokor" (1 Ψ,
ZMHB
).- "
Ungarn
" (1 ɗ, 1 Ψ,
ZMHB
).- "
Hungaria
R. Scholz. | Coll. Döttling" (1 Ψ,
SMNS
).- "
Ungarn
, Izshak, 21.7.[19]93, leg. E. Heise" (1 ɗ, CEH).- "
Ungarn
, Neusiedler See, Fertöuglak, 0 4.05.1999 leg. A. Skale" (1 Ψ,
CAS
).- "
Ungarn
, Prov. Fejer, Umgebung Aba, Handfang
23.V.-7.VI.1997
, leg. F. Bahr" (1 ɗ, 1 Ψ, CHB).- "Ujpest
Ungarn
" (1 ɗ, 2 ΨΨ,
SMTD
).- "
Ungarn
, Halbinsel Tihany, Plattensee 2.6.[19]63, leg. G. Jaeschke" (1 ɗ, 1 Ψ,
DEI
).- "
Ungarn
, Buda-hegy, E.
IV.1969
, leg. Fritzsche" (2 ɗɗ, 4 ΨΨ,
SMTD
).- "
Hungaria
, Cegled, 02.05.[20]02 leg. W. Richter" (2 ɗɗ, 2 ΨΨ, CWR).- "HU Mecseknadasd, Baranya Megye 09.05.[20]03 leg. H. Müller" (1 ɗ,
CHM
).- "
Ungarn
| Kollektion Dr. Fuchs" (1 ɗ, CJS; 1 Ψ,
NSMW
).- "
Ungarn
, A. via, K. Andreas" (1 ɗ,
NSMW
).- "
Ungarn
8.10.03
" (2 ɗɗ.
SMTD
).- "
Ungarn
| Coll. Leonhard" (1 ɗ, 3 ΨΨ,
DEI
).- "Dunakaesz,
Hungaria
, ex Coll. Missionshaus SVD Steyl, Eing. Nr. 3/59" (1 ɗ,
ZMFK
).
Switzerland
:
"Martigny | coll. Stierlin" (3 ɗɗ, 2 ΨΨ,
DEI
).
France
:
"Reims" (1 ɗ, 1 Ψ,
ZMHB
).- "Bretigny s/Orge [Bretigny-sur-Orge, südlich Paris]
9/5/1963
[leg. Bause]" (10 ɗɗ, 9 ΨΨ,
ZMHB
).- "Toul, 6.[19]39" (2 ɗɗ, 4 ΨΨ,
ZMHB
).- "
Frankreich
, Bordeaux 4.[19]43, F. Ermisch lgt." (1 ɗ, 3 ΨΨ,
ZMHB
).- "St. Remy les Chevreuses, 11/5/[19]63" (2 ɗɗ,
ZMHB
).- "Provins (S-et M.)
22 Avr. 1946
Crottin de Cheval" (1 ɗ,
SMNS
).- dto., nur "
11. Mai 47
" (1 Ψ,
SMNS
).- "
Frankreich
St. Denis 6.1974" (1 ɗ, 1 Ψ,
SMNS
).- "Avord-Cher
18.6.1941
Folwaczny" (1 ɗ,
SMNS
).- "
Gallia
/ Ardeche, Vernoux/ Valence, E.V.[19]74, Messulat" (1 Ψ, CJS).- "Etaples |
Frankreich
" (1 ɗ,
SMTD
).- "Metz 5.1916." (1 ɗ, 1 Ψ,
SMTD
).- "Boulogne" (1 ɗ, coll. Andrewes,
NHML
).- "Manche, M. St. Michel,
17.5.64
" (1 Ψ,
NHML
).- "S.
France
: (Var), La Sainte Baume.
16–28.V.1921
. K. G. Blair." (3 ɗɗ, 1 Ψ,
NHML
).- "Environs de Bordeaux, de Vau log er " (1 ɗ, 1 Ψ, coll. Nevinson,
NHML
).- "Bordeaux" (1 ɗ, coll Nevinson,
NHML
).- "Moulins" (1 ɗ, coll. Nevinson,
NHML
).- "Nancy M." (1 ɗ, coll. Andrewes,
NHML
).- "
Gallia
Merid" (1 ɗ, coll. Fry,
NHML
).- "Gall. cent." (1 ɗ, coll. Nevinson,
NHML
).- "
France
" (1 ɗ, coll. Nevinson,
NHML
).- "
Gallia
" (1 ɗ, coll. Fry,
NHML
).- "
Gallia
Mulst" (2 ɗɗ,
SMTD
).- "Indre-et-Loire: Pernay,
2.v.1964
, J.Rabaça legit" (2 ΨΨ, CTB).- "Jura: Arc-Senans, Les Communaux,
20.v.1956
(ex coll. Pasquier)" (1 ɗ, CTB).- "Jura: Les Arsures,
3.vi.1956
(ex coll. Pasquier)" (2 ɗɗ, 1 Ψ, CTB).- "Mayenne: Couptrain, P. Rodary legit" (1 Ψ, CTB).- "Saôneet-Loire: La Selle,
iv.1950
, P.Rodary legit" (1 ɗ, CTB).- "Grasse, A.M. ex Slg. R. Oberthür (Coll. Tondu) Eing. Nr. 4.1956" (1 ɗ, 1 Ψ,
ZMFK
).- "La Salvetat (Hérault), ex Slg. R. Oberthür (Coll. Tondu) Eing.Nr. 4.1956" (1 ɗ,
ZMFK
).- "Lyon, ex Slg. R. Oberthür (Coll. Tondu) Eing.Nr. 4.1956" (1 ɗ,
ZMFK
).- "Wassy, Hte Marne, ex Slg. R. Oberthür (Coll. Tondu) Eing. Nr. 4.1956" (1 ɗ,
ZMFK
).- "Camarque, ex Slg. R. Oberthür (Coll. Tondu) Eing. Nr. 4.1956" (1 Ψ,
ZMFK
).- "Pontavert-France
27.4.1944
, [leg.] Hans Pscher" (1 Ψ,
NMP
).- "Dpt. d’Or,
Ende
Mai 1898
| Riedel" (4 ɗɗ, 3 ΨΨ,
NMP
).- "Pyren. or., Dr. Balthasar” (1 ɗ,
NMP
).
Great Britain
:
"Chesil 1864 J. Gray" (1 ɗ,
SMTD
).- "Hunts: Woodwalton Fen.
6.vi.1938
E. A. Duffy" (1 ɗ,
NHML
).- "W Pontrilas,
29.XI.1929
C. E. T." (1 ɗ, coll. C. E. Tottenham,
NHML
).- "Benfleet
28-iv-1930
C. E. T." (2 ɗɗ, 2 ΨΨ, coll. C. E. Tottenham,
NHML
).- "Bexhill
16.iv.1923
C. E. T." (2 ɗɗ, coll. C. E. Tottenham,
NHML
).- dto., nur "
V.1923
" (2 ΨΨ, coll. C. E. Tottenham,
NHML
).- "Chichester
9.V.1919
C. E. T." (4 ɗɗ, 2 ΨΨ, coll. C. E. Tottenham,
NHML
).- "Warth Dorset or grass
6-VI-37
" (1 Ψ,
NHML
).- "Kent: Dartford, Marshes.
19.IV.1952
. J. F. Burton, Cowdung" (1 Ψ,
NHML
).- "Somerset" (3 ɗɗ, coll. Andrewes,
NHML
).- "Cookham" (3 ɗɗ, 3 ΨΨ,
NHML
).- "
England
. On Cowdung. Totton, Hants,?
Aug. 1953
. SU 355127. G. R. Vardy" (1 Ψ,
NHML
).- "
England
: Dorset (09), Ballard Down, SZ0280,
16.iv.2007
ex horse dung, shaded, coll. D. J.
Mann
, OUMNH-2007-045" (1 ɗ, 1 Ψ,
NHML
).- "Power. Hampstead. 53.7" (2 ɗɗ, coll. W. A. Power,
NHML
).- "Power. Gravesend. 69.20" (1 ɗ, coll. W. A. Power,
NHML
).- "Power. Barnes. 69.18" (1 ɗ,
NHML
).- "Weymouth,
April 21-1901
." (2 ɗɗ, 2 ΨΨ, coll. Donisthorpe,
NHML
).- "Isle of Wight (W Brook), June 5.1899" (1 ɗ, coll. Donisthorpe,
NHML
).- "Rodwell, N. Weymouth,
April 10-1896
" (1 Ψ, coll. Donisthorpe,
NHML
).- "Chingford
VI.1911
" (1 ɗ, coll. H. C. Dollman,
NHML
).- "Etching m-6.24" (1 ɗ, coll. C. J. Saunders,
NHML
).- "Ryde" (1 ɗ, 4 ΨΨ, coll. D. Sharp,
NHML
).- "Edenbridge.
June 1890
" (1 ɗ, coll. D. Sharp,
NHML
).- "Isle of Sheppey. G. C. C." (1 Ψ, coll. G. C. Champion,
NHML
).- "London distr. G. C. C." (2 ɗɗ, 3 ΨΨ, coll. G. C. Champion,
NHML
).- "Abbey Wood. T. W." (1 ɗ, coll. T. Wood,
NHML
).- "Padstow
1.V.1922
S. B." (1 ɗ, 1 Ψ, coll. C. E. Tottenham,
NHML
).- "Barcombe
2.4.27
flood." (1 ɗ, coll. C. J. Saunders,
NHML
).- dto., nur "
24.10.25
" (1 ɗ, coll. C. J. Saunders,
NHML
).- "Brighton
15.5.
38
L. A. Cox" (3 ɗɗ, 1 Ψ, coll. C. J. Saunders,
NHML
).- "
Anglia
" (1 ɗ, 1 Ψ, coll. Fry,
NHML
).
Portugal
:
"P, Ribatejo, UG Santarem, S. de Aire, Kuh, 500 NN,
9.5.1991
, leg. U. Schaffrath" (1 Ψ,
CUS
).- "Monforte: Herdade das Esquilas,
20.v.1997
, A. Zuzarte legit" (1 ɗ, CTB).- "Resende: S. Cipriano,
22.vi.1978
, J. Ramalho legit" (1 Ψ, CTB).- dto., nur "
21.vi.1979
" (5 ɗɗ, 4 ΨΨ, CTB).
Azores:
"Azores | Coll. Felsche, Kauf 20.1918" (2 ΨΨ,
SMTD
).
Spain
:
"29.6.[19]92 (5) Navarra, Valle de Roncal, Garde, Rinderkot, Fery leg." (1 ɗ, 2 ΨΨ, CHF).- "E. Navarra, Sierra de Aralar
1000 m
, 23.5.[19]96 leg. Strke" (2 ΨΨ,
SMNS
).- "E – Navarra, Sierra de Aralar,
700 m
, S. Miguel de Aralar,
23.5.1996
Starke leg." (1 ɗ, 1 Ψ, CJS).- "
Spanien
, Morella, Prov. Castellon de la Plana,
1000 m
, 19./
20.VI.1992
, leg. E. Rößner" (1 ɗ, CER).- "15.6.[19]90
Espana
, Prov. Teruel, S. d.
Albarricin, Orihuela del Tremedal
1500 m
, Schafskot, Fery leg." (1 Ψ, CHF).- "E.-Teruel, Albarracin, 15.-
17.4.2004
, K. Hadulla leg." (1 Ψ, CKHa).- "2.4.[19]88
Espana
, Prov. Madrid, Sierra des Guadarrama, Lozoya, Rinderkot, Fery leg." (1 ɗ, 3 ΨΨ, CHF).- "S[an]. Ildefonso" (1 Ψ,
SMTD
).- "Lisitan. Dyrell." (3 ɗɗ, 1 Ψ, coll. Heyden,
DEI
).- "Serra de Gredos, La Serrada/ Span. VI/
VII.1060
, J. Vives leg." (6 ɗɗ, 3 ΨΨ,
ZSM
).- "Palencia, Pag[...]" (1 ɗ,
ZSM
).- "Sa[ntibáñez]. de Bejar,
Spain
, G. C. C." (1 Ψ, coll. G. C. Champion,
NHML
).- "
Spain
, Avila to El Barraco,
1400 m
(30T359999 4487524) 21-
22.6.2008
" (3 ɗɗ, ex coll. A. Pizzo,
NHML
).
Italy
:
"
4.5.1991
No.
Italien
, Mt. Giorgio bei Savona (
700 m
) leg: Ahrens" (2 ɗɗ, 3 ΨΨ,
CDA
).- "Alp. Mar. piemont., Limone,
1000–1200 m
, Tetti Camila,
20-VI-1951
[leg.] G. Fagel” (1 Ψ,
NMP
).- "
Italien
29/VI 24
, K: 3411" (4 ɗɗ, 1 Ψ,
NMP
).- "16.6.[19]83 Toscana, Chinsi, Schafskot, Fery leg." (1 Ψ, CHF).- "Castel di Sangro, Prov. di Aquila, Paganetti" (1 Ψ,
ZMHB
).- "6.5.[19]90
Italia
, Puglia, Umg. Laterza, S Bari, Schafskot, Fery leg." (1ɗ, CHF).- "Dr. Feige, Roma, Monte Cavo" (2 ΨΨ,
ZMHB
).- "20.6.[19]83 Calabria, Gambarie, Schafskot,
1000 m
, Fery leg." (1 Ψ, CHF).- "27.6.[19]83 Calabria, Sila, Germano
1000 m
, Rinderkot, Fery leg." (1 Ψ, CHF).- "Gargano, Ital./ IV.[19]75, leg. Niehuis" (1 ɗ, CJS).- "
Insel
Elba, Monte Capanne, 9.4.[19]54, Dr. David" (1 Ψ,
ZMHB
).- "Calabrien. Cosenza 6.6.[19]43" (1 Ψ,
SMNS
).- "Calabria – Asprom. Cittanova (RC)
1300 m
, Zomaro,
23.VIII.
[19]81 leg. F. Angelini" (1 Ψ,
SMNS
).- "Liguria, Testico 11.6.[19]
62 W.
Liebmann" (1 ɗ,
SMNS
).- "Südtirol, Staben, Sonnenhang, 20.-23.5.[19]72 Rieger" (1 Ψ,
SMNS
).- "
Italien
, Lazio, Colle
Albani
, Lariano, C. Pacione,
9.IV.1999
lg. Ahrens, Fabrizi" (10 ɗɗ, 5 ΨΨ,
SMTD
).- "
Italia
, Lazio, Colli
Albani
: Vivaro
1 km
E of Nemi,
5.V.1999
leg. S. Fabrizi" (2 ɗɗ, 5 ΨΨ,
SMTD
).- "Lombardia
Italia
" (7 ΨΨ,
SMTD
).- "Calabria, Antonimina 1905, Paganetti" (4 ɗɗ, 2 ΨΨ,
DEI
).- "Calabria, Aspromonte, lg. Paganetti" (1 ɗ,
DEI
).- "San Basilio, Murgien, Paganetti" (2 ΨΨ,
DEI
).- "Apulien, Mte. Gargano" (1 ɗ,
SMNS
).- "Monte Gargano
14.6.1907
legit M. Hilf, Coll.
O
. Leonhard" (1 Ψ,
DEI
).- "Pisa" (1 Ψ,
ZSM
).- "Roma" (1 Ψ,
ZSM
).- "Firenze: M. Giovi,
4.v.1968
, S.Rocchi legit" (3 ɗɗ, 2 ΨΨ, CTB).- "Firenze: Pratolino,
2.v.1970
, S. Rocchi legit" (8 ɗɗ, 11 ΨΨ, CTB).- "Firenze: Vallombrosa, Secchieta,
20.vi.1970
, S. Rocchi legit" (1 Ψ, CTB).- "Ligúria: Savona, Mte. S. Giorgio,
20.iv.1965
, G.Dellacasa legit" (1 ɗ, 2 ΨΨ, CTB).- "Ligúria: Savona, Mte. S. Giorgio,
17.v.1965
, L.Briganti legit" (2 ɗɗ, 1 Ψ, CTB).- "
Italy
, Colle dell’Orso (CB) (E450310, N4607162),
1400 m
09.06.2007
" (2 ɗɗ, ex coll. A. Pizzo,
NHML
).- "
Italy
, Pantano della Zottola, Iserina (E424539, N4619338),
13.06.2007
" (2 ɗɗ, 2 ΨΨ, ex coll. A. Pizzo,
NHML
).- "
Italy
, Lazio: Monti Aurunci, Campodimele env.,
824 m
41°22’30,1’’ N
,
13°32’26,3’’ E
,
31.V.2008
Ahrens & Fabrizi" (1 Ψ,
NHML
).- "
Italy
, Altopiano di Monte: Lago Sefro (MC),
910 m
,
43°07’44,41’’ N
,
12°58’01,10’’ E
,
01.06.2008
" (4 ɗɗ, ex coll. A. Pizzo,
NHML
).- "
Italy
Torino (Ipla) (N400)
23.V.2007
" (2 ɗɗ, ex coll. A. Pizzo,
NHML
).- "Sicilien, Palermo, Ex c. Dr. Studt G." (1 ɗ,
ZMHB
).- "NO-Sizilien, Le Madonie, Piano Zucchi
1000 m
,
23.5.2006
, W. Ziegler" (4 ΨΨ, CWZ).- "I – Sizilien, Madonie, Piano Zucci
22.5.2006
, Heise leg." (2 ɗɗ, 1 Ψ, CEH).- "I – Sizilien, Madonie, Piano Battaglia
23.5.2006
, Heise leg." (1 ɗ, CEH).- "18.4.[19]84 Sicilia, Buccheri, Rinderkot, Fery leg." (1 ɗ, CHF).- "25.3.[19]83 Sicilia, Collesano,
400 m
, Rinderkot, Fery leg." (1 Ψ, CHF).- "25.3.[19]83 Sicilia, Campofehice,
400 m
, Rinderkot, Fery leg." (2 ɗɗ, CHF).- "Sicilien Messina. 25.4.[19]43" (5 ɗɗ, 8 ΨΨ,
SMNS
).- "Sicilien, Messina, 25.4.[19]43, W. Liebmann" (1 Ψ,
DEI
).- "Sicilia
Italia
22.4.[19]33, W. Liebmann" (1 ɗ,
DEI
).- "Sicilia, Randazzo, 3.5.[19]33, W. Liebmann" (11 ɗɗ, 4 ΨΨ,
DEI
).- "Sizilien" (1 Ψ,
SMTD
).- "Sic., Palermo, Dr. Balthasar" (1 Ψ,
NMP
).
Slovenia
:
"Adelsberg [= Postojna] 13.5.[19]09" (2 ɗɗ, 1 Ψ,
SMTD
).- "Marbg. [= Maribor], [leg.] St. Dr. Krause, ex Slg. R. Oberthür (Coll. Tondu) Eing. Nr. 4.1956" (1 Ψ,
ZMFK
).
Croatia
:
"Jug.,
Kroatia
, Istria or., Rabac,
15.–17. v. 1890
Wrase" (2 ΨΨ, CER).- "No 164 Istrien, San Pietro i.S.
4.6.1911
, Schumacher-Spaney S.V." (1 ɗ,
ZMHB
).- "No. 53 Karst, Pared
450 m
, Spaney 29.5.[19]11, Schumacher S.V." (1 ɗ,
ZMHB
).- "No. 123 Karst, Divača-Corgnale
450 m
, 31.5.[19]11, Spaney-Schumacher S.V." (8 ɗɗ, 1 Ψ,
ZMHB
).- "No. 26 Karst, Občina
350 m
.
28.5.[19]11, Spaney-Schumacher S.V." (1 ɗ,
ZMHB
).- "YU / Istrien, Rovin E.5.[19]79 leg. Heidenfelder" (3 ɗɗ, 1 Ψ,
SMNS
).- "Istrien, Pula Umg. 28.5.[19]74 leg. Oschütz" (1 ɗ,
SMNS
).- "Jugosl./ Istrien, Umag, E.3.[19]78" (1 Ψ, CJS).- "19.5.[19]37,
Jugoslavia
, Dalmatien | Crik-Veniza, in Dung | 599 leg. Pause" (1 ɗ,
SMTD
).- "Istria Noghera" (1 Ψ,
SMTD
).- "Meusel,
Croatia
| Coll. Schenkling" (1 ɗ, 1 Ψ,
DEI
).- "
Croatia
, coll. Künnemann" (1 Ψ,
DEI
).- "Istria Rse Ver." (1 ɗ, 1 Ψ, coll. Heyden,
DEI
).- "Fijume [= Rijeka]" (1 Ψ,
ZSM
).- "Istria." (2 ɗɗ,
ZSM
).- "
Kroatien
, Halbinsel Istrien, Rovinj,
14.5.1980
, leg. F. Bahr" (1 ɗ, CLS).- "Pola, Istr., [leg.] Fr. Lang, ex Coll. Missionshaus SVD Steyl, Eing. Nr. 3/59" (1 ɗ, 3 ΨΨ,
ZMFK
).- "Dalm., VI.[19]37, Knin, Bal.-Fr." (1 ɗ,
NMP
).- "Dalmatia" (1 ɗ, 1 Ψ,
NMP
).- "
Kroatien
, Porec,
21.05.1988
, leg. K. Hadulla" (2 ɗɗ, 4 ΨΨ, CKHa).
Bosnia-Herzegovina
:
"
Bosna
, Korična,
O
. Leonhard" (1 ɗ,
DEI
).- "
Bosnia
" (1 ɗ, CJS).
Montenegro
:
"No 451
Montenegro
, Vojnik-Gebirge, Brezovi-do
1000 m
, 27.6.[19]11 Schumacher S.V." (3 ɗɗ, 1 Ψ,
ZMHB
).- "Castelnuovo [= Herceg-Novi], [leg.] Hummler" (1 ɗ,
ZMHB
).- "Dalm. Castelnuovo M. Hilf 1910, Coll.
O
. Leonhard" (1 Ψ,
DEI
).- "Petrovac na Moru,
Montenegro
,
April 1969
, leg. E. Gersdorf" (1 Ψ, CJS).
Serbia
:
"Belgrad
Serbia
" (1 Ψ,
SMTD
).- "Maglič
VIII.1913
,
Montenegro
, legit Jan Obenberger" (1 Ψ,
NMP
).- "Sutorina, Paganetti" (1 Ψ,
NMP
).
Macedonia
:
"YU Maked., Galicica
1600 m
,
20.V.1982
, Krätschmer leg." (1 Ψ, CJS).
Ukraine
:
"Olgino b. Berdjansk, Dnjepr. Gebiet, V/
VI 1937
,
Ukraine
, leg. Nikolajew" (15 ɗɗ, 15 ΨΨ,
ZMHB
).- "
Ukraine
, Dnjeprop. obl., Umg. Berdjansk, leg. Nikolajew,
14.VI.1937
" (1 Ψ,
ZMHB
).- "Umg. Berdjansk, Dnjeprop. obl.,
30.V.1938
, leg. Gulikow" (3 ΨΨ,
ZMHB
).- "
Ukraine
,
24.4.1937
, Mekortj. b. Berdjansk, Dnjepropetr. Gebiet, leg. Nikolajew" (6 ΨΨ,
ZMHB
).- "
Ukraine
, Kamienka,
35 m
s. v. Rowenki [= Rovenki],
VI.1918
, A. Spaney" (1 ɗ, 2 ΨΨ,
ZMHB
).- "Süd-
Russland
, Falzfeinowo a. Dniepr
12.V.–7.VI.1914
, W. Ramme S.G." (5 ɗɗ, 1 Ψ,
ZMHB
).- "
Ukraine
,
9.6.1935
, Chomutowsk. Steppe, b. Mariupol, leg. Porischenko" (1 ɗ,
ZMHB
).- "Krim, Simferopol,
24.V.1942
, Haltenorth S.G." (1 ɗ, 1 Ψ,
ZMHB
).- "Odessa" (1 ɗ,
SMTD
).- "Crimea" (2 ɗɗ,
NHML
).- "Sebastopol Krim
10.4.1903
[leg.] W. Pligiuski" (1 Ψ,
NMP
).
Russia
:
"Rostov na Donu, R. m.” (1 Ψ,
NMP
).- "Südrussland, Sarepta | H. Rolle Berlin | ex coll. Hopffgarten Mülverstedt." (1 Ψ,
NMP
).- "Terskaja obl., St. Mineralni Wod.,
30.IV.1928
" (13 ɗɗ, 1 Ψ,
ZMHB
).- "Dongebiet,
28.V.–3.VI.1918
, A. Spaney" (1 Ψ,
ZMHB
).- "S.
O
.
Russland
, Tschernyj-Jar, Wolga, Kracht S.G." (1ɗ,
ZMHB
).- "Daghestan (
Rossia
), Nowyi Birjusjak, leg. Worobjew, 6.6.[19]57" (1 ɗ,
ZMHB
).- "S.E.
Russia
, Volsk district. R. S. Kozlov. B. M.
1931- 39
" (2 ɗɗ, 2 ΨΨ,
NHML
).- "Transcaspia, Neu-Saratow" (1 Ψ,
NHML
).- "Kaukas. Schneider" (1 Ψ,
SMTD
).- "Caucasus, Alexandrofska, Ryssel S.V. / 18.4.[19]01" (3 ɗɗ,
ZMHB
).- "Caucasus" (1 Ψ,
NHML
; 1 ɗ,
SMNS
).
Kazakhstan
:
"Indersk." (2 ɗɗ, 1 Ψ, coll. E.
Y
. Western,
NHML
).
Romania
:
"Desert. Deliblát, Banat, Holtz" (4 ΨΨ,
ZMHB
).- "
Rumänien
, Donaudelta, Unirea [bei Brăila],
11.VI.1988
leg. Hornburg" (1 ɗ, 3 ΨΨ, COH).- "
Roumanie
, Comana Vlasca, A. L. Montandon" (1 ɗ,
DEI
).- "Dobroudja. Mangalia. A. L. Montandon" (1 ɗ,
DEI
).- "Jassy [= Iasi]" (3 ɗɗ,
ZSM
).- "Bukarest" (1 ɗ,
ZSM
).
Bulgaria
:
"Mandra,
Bulgarien
,
5.5.1933
, leg. Fuss" (1 ɗ,
NMP
).- "
Bulgarien
, Kitschevo, N Varna
28.V.1995
, leg. H. Kalz" (1 Ψ, CHKa).- "BG, Sveti Vlas,
16.V.2005
, leg. H. Müller" (3 ɗɗ, 6 ΨΨ,
CHM
).- "
Bulgaria
, Sveti Vlas, 08.05.- 19.05.[20]05 leg. W. Richter" (6 ɗɗ, 2 ΨΨ, CWR).- "
Bulgarien
, Umg. Sozopol, 9.6.-
20.6.1978
, leg. Bringmann" (1 Ψ, CER).- "SO-Bulgarien, südl. Sosopol, Umg. Kavacite,
30. Mai 1982
, leg. H. Wendt" (1 ɗ,
ZMHB
).- "SO-Bulgarien, südl. Burgas, Umg. Acheloi, 1./
2.VI.1982
, leg. H. Wendt" (1 Ψ,
ZMHB
).- "
Bulgarien
, U. Nessebar, 4.6.[19]67, K. Ermisch leg." (1 Ψ,
ZMHB
).- "
Bulgarien
, Primorsko, 12.6.[19]85, leg. Kriska" (1ɗ, CER).- "SO-Bulgarien: Zarewo (= Mičurin),
13.V.2002
, leg. E. Rößner" (1 Ψ, CER).- "SO-
Bulgarien
, Strandža-Gebirge: Malko Tarnovo,
14.V.2002
, leg. E. Rößner" (1ɗ, CER).- "Sofia" (1 Ψ,
SMTD
).- "
Bulgaria
,
55 km
SE Burgas, Lozenec
13.V.2002
, leg. L. Schmidt" (1 ɗ, 1 Ψ, CLS).- "
Bulgaria
,
35 km
NW Burgas,
5 km
W Ajtos, Trockenhang,
18.V.2002
, leg. L. Schmidt" (1 ɗ, 1 Ψ, CLS).- "
Bulgaria
,
37 km
SE Burgas, Jasna poljana, Viehweide,
17.V.2002
, l. L. Schmidt" (2 ɗɗ, CLS).- "Mandra,
Bulgaria
,
20.5.1933
, Fuss leg." (1 ɗ, 4 ΨΨ,
ZMFK
).
Greece
:
"
Graecia
, L. Salvator." (1 ɗ,
NMP
).- "Ossa,
Graecia
" (1 ɗ, 1 Ψ,
NMP
).- "
Greece
(C.
Macedonia
)
258 m
, Lake Kerkini env., Akitoschori N 41°16’72,5’’/E 023°10’15,3’’,
15.IV.2003
leg.
O
. Hillert" (1 ɗ, 1 Ψ, COH).- "
Griechenland
, Prionia Gebirge, Olymp,
1000 m
,
27.VI.- 17.VII.1995
, leg. T. Kriska" (1 ɗ, CER).- "12.4.[19]
84 N.
Griech. westl. Kavalla,
300 m
, Schafskot, Fery leg." (1 Ψ, CHF).- "13.4.[19]84 Griech., Larissa, Schafskot, Fery leg." (1 ɗ, CHF).- GR,
Makedonien
, Nomós Kavála,
10 km
N Pale á Kavála,
600 m
, Macchie + Kastanienwald
20.IV.1994
, leg. Schawaller" (1 Ψ,
SMNS
).- "GR, Ossa, Oros,
500 m
, Stomio,
27.5.1990
, Persohn leg." (1 ɗ,
SMNS
).- "GR – Thessalie, Stomio
5.7.2006
, Heise leg." (1 ɗ, CEH).- "GR;
5 km
ö Kato Vermion,
1000 m
, Baehr 1.5.[19]83" (2 ΨΨ, CJS).- "GR Vermion Oros, Kato Vermion
1300 m
, Krätschmer lg.
IV.1977
" (1 ɗ, CJS).- "
Graecia
/ Thessalia, Ossa Oros/ Stomio
1000 m
/
27.V.1990
, Persohn leg." (1 ɗ, 1 Ψ, CJS).- "GR:
10 km
w. Veria.
700 m
.
Baehr 30.4.[19]83" (1 ɗ, CJS).- "
Greece
/ Monodéndri, Vikou-Gebirge
03.–06.V.1993
leg. Ringel" (1 Ψ,
CHR
).– "GR, Peloponnes,
1400 m
, Exohori, Prov. Messinia,
2.7.2005
, leg. W. Ziegler" (2 ɗɗ, 2 ΨΨ, CWZ).- "
Griechenland
, Peloponnesos: Araxos (
30 km
w Pátrai),
26.–31.V.1993
, leg. E. Rößner" (1ɗ, CER).- "Balkan, Corfu. Paganetti [19]03" (1 ɗ,
DEI
).- "16.4.[19]84 Corfu, Roda, Pferdekot, Fery leg." (1 ɗ, 1 Ψ, CHF).- "Rhodes" (1 ɗ,
NMP
).
Turkey
:
"Europ.
Türkei
: Istranca-dag bei Demirköy, Paß,
700 m
, Heinz leg.
2.V.1973
" (1 Ψ, CJS).- "Constantinopel
V.1898
, W. Gottwald S.V." (1 ɗ,
ZMHB
).- "Kleinasien, Biga, 22.4.[19]16, Dr. Bauer S.G." (1 ɗ,
ZMHB
).- "
Asia
minor
, Biledjek, v. Bodemeyer" (1ɗ,
ZMHB
).- "
Angora
. Asia min., V. M. Duchon" (4 ɗɗ, 2 ΨΨ,
ZMHB
).- "
Türkei
(Centr.), Puturgi bei Malatya,
10.VI.1993
, leg. Kriska" (1 ɗ, COH).- "
Turcia
m., Prov. Antalya,
20 km
NE Demirtas, Hochebene
1400 m
,
36°31’32’’ N
,
32°23’26’’ E
,
28.V.2006
, leg. A Kopetz" (2 ΨΨ, CAK).- "
Türkei
, Anatolien, Nohutlu (
90 km
sö von Malatya), ca.
1700 m
,
5.– 9.VI.1993
, leg. A. Schröder" (3 ɗɗ, 3 ΨΨ, CER).- "Ararat, Wagn[er]." (1 ɗ,
ZMHB
).- "Tauria, Part. | 26959" (1 ɗ,
ZMHB
).- "Kleinasien, Anatolien | Kollektion Dr. Fuchs" (1 Ψ, CJS).- "
O
.
Türkei
20.–26.VI.
[19]75, 30–
40 km
ö Bingöl, leg. D. Berhauer" (1 Ψ, CJS).- "Kleinasien, Anatolien | Kollektion Dr. Fuchs" (3 ɗɗ, 3 ΨΨ,
NSMW
).- "v. Bodemeyer, Klein-Asien, Eski-Chehir [= Eskişehir] | Kollektion Dr. Fuchs" (1 ɗ,
NSMW
).- "
Asia
minor
, Burna, v. Bodemeyer | Kollektion Dr. Fuchs" (1 Ψ,
NSMW
).- "
Asia
minor
, Sultan Dagh, v. Bodemeyer" (1 ɗ,
NMP
).- "Klein-Asien, Eski-Chehir, v. Bodemeyer" (2 ɗɗ,
NMP
).- "Eski Chehir, Coll. Purkynĕ." (1 ɗ,
NMP
).- "
Asia
minor
, Ak-Chehir, v. Bodemeyer” (1 ɗ, 1 Ψ,
NMP
).- "v. Bodemeyer,
Anatolia
,
Asia
minor
” (1 ɗ,
NMP
).- "
Anatolia
, Asie Mineure, Coll. Obenberger” (3 ɗɗ, 3 ΨΨ,
NMP
).- "v. Bodemeyer, Klein Asien, Bulghar Maaden” (1 ɗ, 1 Ψ,
NMP
).
Georgia
:
"Borshom Caucasus,
O
. Schneider" (1 Ψ, coll. Heyden,
DEI
).
Iran
:
"N.
Iran
2.VI.
[19]77, 40 km s Sari | Elb., leg. D. Bernhauer" (1 ɗ, CJS).- "
Persia
Astrabad [= Gorgan]" (4 ɗɗ, 4 ΨΨ,
SMTD
).- "N-Iran, Prov. Teheran, C-Elburz, Damavand Mt. Gosfand Sara, Larijan,
35° 52’ N
52° 07’ E
,
2300 m
,
29.05.2008
, leg. A. Skale (11a)" (4 ɗɗ, 3 ΨΨ,
CAS
).- "Elbrus Geb.,
Persien
, Rttr." (2 ɗɗ,
NMP
).- "
Persien
, Luristan, v. Bodemeyer" (2 ɗɗ,
NMP
).
Azerbaijan
:
"Lenkoran, Dr. Martin, ex Slg. R. Oberthür (Coll. Tondu) Eing. Nr. 4.1956" (2 ΨΨ,
ZMFK
).
Not localised:
"L. Chamboy,
8.8.38
" (1ɗ,
ZMHB
).- "Transcaspien | Coll. Döttling" (1 Ψ,
SMNS
).- "Wan
22.4.12
" (1 ɗ, 1 Ψ,
ZSM
).- "Kasp. Meer-Geb., Petrowsk, 1897 Korb" (1 ɗ,
ZSM
).
Redescription.
Body length:
8–13 mm
, body oval, weakly convex, dull; head and pronotum black with dark green metallic sheen, rarely cupreous or intensively metallic green; elytra yellow with rows of asymmetric dark spots sometimes arranged in stripes; ventral face, pygidium and legs black with very weak metallic sheen, rarely cupreous; dorsal pilosity short and light.
Head. Antenna brown, clavus black; anterior edge of clypeus reflexed, with transverse wrinkles, head coarsely punctate to granulate between frontal carina and vertex.
Pronotum. Lateral margins behind anterior angles straight or convex, anterior angles weakly rounded, posterior angles sinuate, basis with complete and distinct marginal line (Fig. 2AJ); punctation coarse, dense, very dense and granulate beside anterior angles, on midline and beside posterior angles with narrow smooth impunctate sections.
Elytra. Yellow, base narrowly dark, sutural interval widely or completely dark with green sheen, epipleura completely light; spots black without or with only very weak green sheen, asymmetric (Fig.
2I
, 2J), base always without spots, spots mostly confluent (Figs. 2K–M), sometimes condensed into symmetric stripes (Figs. 2N, 2
O
), or covering the intervals almost entirely (Fig. 2
O
); intervals chagreened, punctation even, granulate; striae shallow, weakly punctate, 7th stria weakly curved.
Pygidium. Base with marginal line, dull, punctation fine, moderately dense and simple, superficial.
Aedeagus. Figs. 3A, 3B, 3Q–AB; lamella copulatrix: 3AG–AK, 3AP–AT (see
Table 1
).
Remarks.
We searched for the collection J. G. Kugelann (
1753–1815
) to find the
type
specimens of his
Copris medius
. According to
Horn et al. (1990
: 218) his beetle collection was passed to Hoffmann at Warsaw. The collection Kugelann is currently preserved at the Museum and Institute of Zoology, Polish Academy of Sciences, Warsaw. According to information of the curator, Mrs. D. Mierzwa (personal communication, 2008), there is no specimen in the collection under either the name '
Copris
[
Onthophagus
]
medius
' or
O
.
vacca
. Consequently it must be assumed that the
type
material was lost or destroyed. For an unambiguous definition of
O
. medius
and in order to differenciate between the two European species of the
O
. vacca
complex, we designate here a
neotype
for
O
. medius
. We chose for
neotype
a male from
Poland
(Lyck = Ełk).
Male
,
major form
: Reflexed portion of anterior edge of clypeus bluntly truncate, rarely weakly sinuated (Fig. 2Q); frontal carina absent, impunctate on this spot; carina of vertex with large lamella with 1st sides basally parallel, apically with a sharply pointed horn being almost rectagular erect and with s-shape arch (lateral view) (Fig. 2AF–AG); Pronotum on anterior declivity smooth, medially with a deep impression, marginal line of base beside posterior angles complete (Fig. 2AJ).
Male
,
minor
form
: Anterior edge of clypeus sinuate (Fig. 2S); frontal carina weakly indicated to slightly elevate, curved; carina of vertex with a small lamella being slightly elevated at middle; pronotum anteriorly with rugose punctures, medially with a pair of small, smooth convexities, marginal line of base beside posterior angles complete.
Female,
major form
: Anterior edge of clypeus weakly sinuate; frontal carina distinct, elevate, weakly curved; carina of vertex low, as wide as entire posterior edge of head, each end blunt and weakly elevated (Figs.
2X
, 2AC); pronotum at anterior marginal declivity widely smooth and only sparsely punctate, medially with a slightly anteriorly produced and medially slightly incised convexity, beside it on each side with a weak convexity, marginal line of base beside posterior angles complete (Fig. 2AJ).
Female,
minor
form
: Anterior edge of clypeus weakly sinuate; frontal carina short, elevate; carina of vertex wide and low, without or only with slightly indicated elevated ends (Figs. 2
Y
, 2AE); pronotum anteriorly with rugose punctures, medially with a slightly transeverse smooth convexity, basal marginal line complete.