Dung beetles (Scarabaeidae, Scarabaeinae) of the Foothills-Andean Forest strip of Villavicencio, Colombia Author Cárdenas-Bautista, Johann Stephens Author Parada-Alfonso, Jenny Andrea Author Carvajal‑Cogollo, Juan E. text Check List 2020 2020-07-03 16 4 821 839 http://dx.doi.org/10.15560/16.4.821 journal article 10.15560/16.4.821 1809-127X Dichotomius mamillatus (Felsche, 1910) Materials examined. COLOMBIA • 1♀ , 23.74 mm (mean); Meta , Villavicencio , Vereda El Carmen , “La Loma” Farm ; 04°08′28.18″N , 073°40′01.90″W ; 730 m a.s.l. ; 18 Oct. 2018 ; Cárdenas Johann and Parada Jenny leg.; wooded pasture; Crpat 3; UPTC-In-00054 . • 1♂ , 23.74 mm (mean); Meta , Villavicencio , Vereda El Carmen , Caño Blanco–Caño Buque ; 04°08′21.85″N , 073°40′12.19″W ; 774 m a.s.l. ; 17 Oct. 2018 ; Cárdenas Johann and Parada Jenny leg.; riparian forest; Crbrt 4; UPTC-In-00055 . Identification. This species can be distinguished from other Dichotomius species by the head with transverse roughness in the clypeus, gena and frons, the presence of a short, thick and blunt horn at the height of the gena with two small horns adjacent to the base of the central horn; pronotum with two small protuberances in the anterior medial region ( Sarmiento-Gárces and Amat-García 2014 ). Distribution. The species is distributed in the Colombian and Ecuadorian Amazon, where it is a very wellrepresented species, extending to the foothills up to 1300 m a.s.l. ( Sarmiento-Gárces and Amat-García 2009 ; Sarmiento-Gárces and Amat-García 2014 ). In the study area, it was observed between 750–850 m a.s.l. and seen in the habitats of riparian forest, high secondary forest, and wooded grassland, although it was rare species (59 individuals).