Morphology-based cladistics splinters the century-old dichotomy of the pied harvestmen (Arachnida: Gonyleptoidea: Cosmetidae) Author Medrano, Miguel Author Kury, Adriano Brilhante Author Mendes, Amanda Cruz text Zoological Journal of the Linnean Society 2022 2021-08-30 195 585 672 journal article 113070 10.1093/zoolinnean/zlab043 e070d52f-d008-4f08-988a-a8b17c27418f 0024-4082 6959516 1C89556A-12CB-43B7-9B49-E02EFF1543D3 PROTUS SIMON, 1879 Protus Simon, 1879: 193 . Type species: Protus insolens Simon, 1879 , by monotypy. Etymology: Protus is a character of Plutarch´s Dion. Gender masculine. Diagnosis: It differs from the other Discosomaticinae by having two white tubercles in area III ( Fig. 31B, C ) and by having the dorsal process of glans slender and acuminate ( Figs 35G , 41A ). Some species have the facies of Sibambea and Kayania because of the coloration of coxa IV with yellow spots ( Figs 33B , 44C ), for having a yellowish border in anterior margin of DS ( Figs 25 , 37 , 42A ) and for the absence of constriction I in the outline of DS in dorsal view. It differs from Sibambea by having a long VP of penis ( Fig. 35G , except for P. distinctus and P. bolivari ). Protus differs from Kayania and Marronia by not having strong ornamentation on the scutal area III ( Fig. 35F vs. Fig. 24 ). Figure 30. Map of known distribution of Protus . Numbers correspond to the following WWF ecoregions: 1 = Ucayali moist forest, 2 = Iquitos varzea, 3 = Southwest Amazon moist forest, 4 = Solimões-Japurá moist forest, 5 = Juruá-Purus moist forest, 6 = Madeira-Tapajos moist forest, 7 = Japurá-Solimões-Negro moist forest, 8 = Uatuma-Trombetas moist forest, 9 = Mato Grosso seasonal forest. Included species: Protus bolivari ( Roewer, 1952 ) comb. nov . , P. distinctus ( Avram & Soares, 1983 ) comb. nov . , P. insolens Simon, 1879 , P. quadripunctatus ( Roewer, 1947 ) comb. nov . , P. marllusi Medrano, Kury & Mendes sp. nov . and P. speciosus ( Roewer, 1928 ) comb. nov . Combined distribution: Upper Amazon basin, eastern Andean slope in Peru ( Fig. 30 ).