Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae)
Author
Selnekovič, Dávid
D930D757-D929-4248-A29C-A857E0505345
Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.
david.selnekovic@uniba.sk
Author
Jäch, Manfred A.
D749707A-8823-4110-8D51-BF5AF4E2820F
Naturhistorisches Museum Wien, Burgring 7, A- 1010 Wien, Austria.
manfred.jaech@nhm-wien.ac.at
Author
Kodada, Ján
6E88BFBB-8769-44EC-8285-29E357CEC064
Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.
jan.kodada@uniba.sk
text
European Journal of Taxonomy
2024
2024-09-11
957
1
229
https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2651/12247
journal article
10.5852/ejt.2024.957.2651
2118-9773
13773385
9957F2DF-3F31-4B58-84BC-2F8E0F45C6E3
Elmomorphus glabriclunis
sp. nov.
urn:lsid:zoobank.org:act:
97854318-7333-4566-90A6-D1CADF579AE4
Figs 100–101
,
106C
,
115E
Differential diagnosis
Elmomorphus glabriclunis
sp. nov.
(
Fig. 100
) is characterised by having a strongly convex body and the dorsal plastron confined only to small areas around the antennal insertions. Elytral striae are absent, and ventrite 1 is entirely covered with plastron. This species strongly resembles
E. simplex
sp. nov.
and
E. comosiclunis
sp. nov.
, but it differs from both in the ventrite 5, which is strongly convex anteriorly and deflexed posteriorly with a sizeable median area without plastron. In
E. simplex
, ventrite 5 is evenly convex, and the area lacking plastron along the posterior margin is narrow. In
E. comosiclunis
, ventrite 5 is simple and evenly convex in females, while males have a large flat oval area with remarkably long setae.
Etymology
The epithet, a noun (nominative singular) in apposition, is a combination of the Latin words ‘glaber’ (‘glabrous’) and ‘clunis’ (‘rump’). It refers to the glabrous area on ventrite 5.
Type material
Holotype
CHINA
–
Hong Kong
•
♂
; “
HONGKONG
leg.
Dudgeon
[printed] | Tai Po Kau
19.1.1978
K [handwritten by
M.A. Jäch
]”;
NMW
.
Paratypes
CHINA
–
Hong Kong
•
2 ex.
; “HONGKONG
1977-1979
Benthic Coll.
Tai Po Kau
leg.
Dudgeon
”;
NMW
•
1 ♂
; “
HONGKONG
Tai Po Kau
2.11.1978
leg.
Dudgeon
(
K
)”;
NMW
•
1 ex.
; “
Tai Po Kau
riv.
27.IV.1978
K5 |
HONGKONG
leg.
Dudgeon
”;
NMW
•
1 ex.
; “
HONGKONG
Tai Po Kau
9.3.1978
leg.
Dudgeon
(
K
)”;
NMW
•
1 ex.
; “
HONGKONG
Tai Po Kau
2.1.1979
leg.
Dudgeon
(3)”;
NMW
•
1 ex.
; “
HONGKONG
Tai Po Kau
28.7.1983
leg.
Dudgeon
(upstr.)”;
NMW
•
1 ex.
; “
HONGKONG
,
Tai Po Kau
12.-
13.4.1984
leg.
Dudgeon
(contr.)”;
NMW
•
2 ex.
; “
HONGKONG
(3) [CWBS 4]
1992
N.T.-
Plover C.
[ove] Res.[ervoire]
25.VI.
leg.
Jäch
”;
NMW
•
1 ♂
; “
HONGKONG
(7) [CWBS 8] 1992 N.T.-
Tai Po N.Res.
27.VI.
leg. Jäch”;
NMW
.
–
Fujian Province
•
1 ex.
; “
CHINA
:
FUJIAN
,
Yong’an
,
5 km
SW Xiyang
,
550 m
Ziyungdong Shan
,
25. 1. 1997
, leg.
Ji
&
Wang
(
CWBS
256)”;
NMW
•
1 ♂
,
1 ♀
; “
CHINA
:
FUJIAN
,
Longyan
Jiangshan (
20 km
N
Longyan
)
Keshan
monastery,
900 m
Meihua Shan
,
28.1.1997
leg.
H. Schönmann
(
CWBS
260)”;
NMW
•
3 ♂♂
,
6 ♀♀
; “
CHINA
:
FUJIAN
,
Longyan
Shizong, Shangfang Shan
850m
31.1.1997
leg.
Ji
&
Wang
(
CWBS
264)”;
CKB
,
NMW
•
5 ♂♂
,
3 ♀♀
; “
CHINA
:
Fujian
,
Yong’an
5 km
N Xiyang
,
400m
26.1.1997
leg.
Ji
&
Wang
(
CWBS
258)”;
CKB
,
NMW
.
–
Guangdong Province
•
1 ex.
; “
CHINA
:
Guangdong
Prov.
50 km
E Fengkai
23°27′36″N
111°54′36″E
31.10./
2.11.2001
, ca.
150 m
Jäch
&
Komarek
(
CWBS
455)”;
NMW
•
1 ♂
,
4 ♀♀
; “
CHINA
:
Guangdong
Prov.
50 km
E Fengkai
23°27′04″N
111°53′53″E
1.11.2001
, ca.
300-400 m
Jäch
&
Komarek
(
CWBS
456)”;
NMW
•
1 ex.
; “
CHINA
:
Guangdong
Prov.
60 km
E Fengkai
23°26′38″N
111°58′10″E
1.11.2001
, ca.
230 m
Jäch
&
Komarek
(
CWBS
458)”;
NMW
•
2 ♂♂
,
1 ♀
; “
CHINA
:
Guangdong
Prov.
45 km
N Zengcheng
23°37′28″N
113°50′10″E
13.11.2001
, ca.
500 m
leg.
M. Wang
(
CWBS
495)”;
NMW
•
1 ♂
,
1 ♀
; “
CHINA
:
GUANGDONG
Prov.
W of Qixing
,
1–3.v.2011
HEISHIDING nature reserve (forested stream valley; at light)
23°27.9′N
,
111°54.3′E
,
190 m
M. Fikáček
&
J. Hájek
leg.”;
NMPC
.
–
Guangxi
Autonomous Region
•
8 ex.
; “
CHINA
: SE-Guangxi,
Distr. Yulin
,
Liuwan Mts.
SW
Yulin
|
18.11.1993
300m
leg.
Schönmann
(23) [CWBS 45]”;
CKB
,
NMW
•
2 ex.
; same collection data as for preceding, but “leg.
Schillhammer
”;
NMW
•
1 ex.
; same collection data as for preceding, but “leg. L. JI”;
NMW
•
11 ♂♂
,
11 ♀♀
; “
CHINA
SE-Guangxi
Distr. Yulin Liuwan Mts.
SW
Yulin
|
20.11.1993
120m
(26) [CWBS 49] leg.
Schönmann
”;
CKB
,
NMW
•
7 ex.
; “
CHINA
,
SE-Guangxi
Distr. Yulin Liuwan Mts.
SW
Yulin
|
17.11.1993
Kuishan
,
600-700m
leg.
L. Ji
(21) [CWBS 43]”;
NMW
•
2 ex.
; same collection data as for preceding, but “leg.
Schillhammer
”;
CKB
,
NMW
•
5 ex.
; same collection data as for preceding, but “leg.
L. JI
”;
NMW
•
7 ex.
; “
CHINA
SE-Guangxi
Distr. Yulin Liuwan Mts.
SW
Yulin
|
16.11.1993
350-400m
leg.
Schönmann
(20) [CWBS 42]”;
NMW
•
3 ex.
: same collection data as for preceding, but “leg.
Schillhammer
”;
NMW
•
1 ♂
,
2 ♀♀
; “
CHINA
SE-Guangxi
Distr. Yulin Liuwan Mts.
SW
Yulin
|
18.11.1993
200m
, (24) [CWBS 46] leg.
Schönmann
”;
NMW
•
1 ♀
: “
CHINA
SE-Guangxi
Distr. Yulin Liuwan Mts.
SW
Yulin
|
17.11.1993
Kuishan
,
300m
leg.
Schönmann
(22) [CWBS 44]”;
NMW
.
Fig. 100.
Elmomorphus glabriclunis
sp. nov.
, paratype, male from China, locality CWBS 49 (CKB), TL: 3.39 mm.
Type locality
China
,
Hong Kong Special Administrative Region
, New Territories, Tai Po Kau Nature Reserve.
Description
Measurements (mm): TL:
♂♂
3.14–3.63 (3.39± 0.11, n=15),
♀♀
3.36–3.70 (3.50 ±0.12, n=10); PL:
♂♂
0.67–0.81 (0.73 ± 0.04, n= 15),
♀♀
0.69–0.81 (0.73 ±0.03, n=10); PW:
♂♂
1.52–1.69 (1.60± 0.05, n= 15),
♀♀
1.52–1.79 (1.65± 0.08, n =10); EL:
♂♂
2.47–2.93 (2.66 ± 0.10, n=15),
♀♀
2.63–2.97 (2.77 ± 0.11, n =10); EW:
♂♂
1.79–2.02 (1.90± 0.06, n =15),
♀♀
1.80–2.02 (1.95± 0.08, n =10); PhL: 0.53–0.61 (0.56± 0.02, n =15); PrL: 0.40–0.48 (0.43± 0.02, n= 15).
Body oval, moderately convex dorsally (
Fig. 101
). Integument black; mouthparts, antennae, and legs reddish brown. Pubescence yellowish. Plastron on dorsum confined to an area around antennal insertions; ventral plastron present on entire surface, except on prosternal process, median part of metaventrite, and median part of terminal ventrite.
Head smooth with round punctures slightly smaller than an eye facet, puncture distance 0.5–1.0 × diameter. Labrum microreticulate on exposed portion, with small setiferous punctures; setae equal in length in both sexes; anterior margin finely emarginate. Eyes oval, interfacetal setae short. Antennae 10-segmented, densely setose.
Pronotum moderately convex; PW/PL:
♂♂
2.06–2.34 (2.21± 0.09, n=15),
♀♀
2.10–2.46 (2.26 ± 0.13, n= 10); rim of anterior margin as wide as 2–3 eye facets, interrupted in middle; anterior angles strongly deflexed, prominent; lateral pronotal sides convergent, rounded; surface smooth, with round setiferous punctures. Prosternal process with lateral edges straight, posterior edge rounded, lateral portions moderately raised, without groups of long setae, median keel arcuate. Scutellum wider than long, smooth, with minute punctures. Median part of metaventrite slightly convex, posteriorly widened; groups of long setae absent in both sexes; lateral margins of metaventral process raised. Elytra oval, moderately convex, widest before middle, EL/EW:
♂♂
1.30–1.50 (1.40± 0.06, n =15),
♀♀
1.38–1.48 (1.42± 0.04, n= 10); surface microreticulate, with scattered round punctures. Tibiae slightly curved, protibia ca 1.6× as long as protarsus; PrTL/PL:
♂♂
1.02–1.19 (1.11 ± 0.05, n=15),
♀♀
0.96–1.21 (1.09± 0.08, n =10). Terminal protarsomere as long as three preceding tarsomeres combined.
Abdomen covered with plastron, except on sizeable oval area on ventrite 5. Ventrite 1 with two longitudinal keels medially. Ventrite 5 with short longitudinal keel before apex in females. Aedeagus (
Fig. 101
): phallobase slightly expanded basally, PhL/PrL: 1.13–1.51 (1.29± 0.09, n= 14); parameres rather long and narrow, curved ventrad in apical half, apices rounded (lateral aspect); penis apically narrowly rounded; sclerotised fibula slender. Bursa copulatrix (
Fig. 106C
) with several microsclerites located laterally in proximal portion.
Secondary sexual dimorphism
Ventrite
5 in
females with short longitudinal keel before apex.
Distribution
China
(
Fujian
,
Guangdong
,
Guangxi
,
Hong Kong
) (
Fig. 115E
).