Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae) Author Pall-Gergely, Barna https://orcid.org/0000-0002-6167-7221 Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary pallgergely2@gmail.com Author Hunyadi, Andras Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary Author Vermeulen, Jaap J. JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands Author Grego, Jozef Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia Author Sutcharit, Chirasak Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand Author Reischuetz, Alexander Puechhaimgasse 52, A- 3580 Horn, Austria Author Dumrongrojwattana, Pongrat Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand Author Botta-Dukat, Zoltan Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary Author Oerstan, Aydin 12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA Author Fekete, Judit University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary & Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary Author Jochum, Adrienne https://orcid.org/0000-0002-6624-6412 Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland & Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany text ZooKeys 2023 2023-02-13 1147 1 177 http://dx.doi.org/10.3897/zookeys.1147.93824 journal article http://dx.doi.org/10.3897/zookeys.1147.93824 1313-2970-1147-1 9BB9881B0076473D8E53155D37CA1F50 FF2B6B317B505F9EA0E1000BDCD16CE7 Angustopila gracilis Pall-Gergely & Hunyadi sp. nov. Fig. 65 Type material. Holotype : Laos • 1 empty shell (H: 0.94 mm, D: 0.85 mm); Bolikhamsai Province, 15 km southeast from centre of Lak Sao, 4.5 km northeast on a side road, Phu Phako, limestone gorge (locality code: 2019/110); 18°06.55'N , 105°03.78'E ; 510 m a.s.l.; 2 Oct. 2019; A. Hunyadi leg.; HNHM 105296. Paratypes : Laos • 699 adult + 207 juvenile shells; same data as for the holotype; coll. HA (juvenile shells are also of characteristic shape for the species, and thus, they are also selected to be paratypes) • 10 shells, same data as for the holotype; coll. JJV. Diagnosis. A medium-sized, concave-conical Angustopila species with conspicuously narrow initial whorls and wide body whorl, aperture wide with a regularly developed parietal tooth, and a very weak, blister-like lower palatal tooth. Description. Shell of normal size for the genus, higher than wide or rarely slightly wider than high; off-white, concave-conical with conspicuously narrow initial whorls and wide body whorl; last whorl widest from standard apertural view; protoconch consists of 1.25 whorls, spirally striated with the exception of the first protoconch whorl; teleoconch with weak, irregular radial ribs and equidistantly, sparsely-spaced, spiral striae (ca. 12-16 on body whorl from standard apertural view); whorls 4.25-4.5, rounded; aperture oblique to shell axis in lateral view; umbilicus wide, slightly excentric; aperture nearly rounded, wide; sinulus wide, weakly separated; peristome expanded, not reflected; parietal callus not detached from penultimate whorl; parietal tooth elevated, straight, starts in some distance from peristome, rather long; lower palatal tooth tiny, blister-like. Figure 65. Angustopila gracilis Pall-Gergely & Hunyadi, sp. nov. A paratype B-I holotype (HNHM 105296). Apertural ( A, B ), lateral ( C ), apical ( D ) and ventral ( E ) sides of the shell; aperture ( F ); sculpture on the protoconch ( G ), ventral ( H ) and frontal ( I ) surface of the body whorl. Measurements (in mm). H = 0.89-0.97, D = 0.82-0.95, H/D*100 = 93.7-113.4 ( n = 7), RUD = 26.5-31.8 ( n = 4). Differential diagnosis. Angustopila antidomedon sp. nov. and A. pustulata sp. nov. possess a less rounded (rather depressed-rounded) body whorl and a less rounded aperture. Moreover, none of those species have such slender initial whorls. Etymology. This species is named after the slender (Latin: Angustopila gracilis ) initial whorls. Distribution. Angustopila gracilis sp. nov. is known from the type locality only in Bolikhamsai Province of Laos (Fig. 31 ).