Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo Author Liew, Thor-Seng Author Schilthuizen, Menno Author Vermeulen, Jaap Jan text Zoological Journal of the Linnean Society 2009 2009-10-26 157 3 515 550 https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x journal article 10.1111/j.1096-3642.2009.00526.x 0024-4082 5443023 EVERETTIA KLEMMANTANICA GUDE, 1918 Everettia klemmantanica Gude, 1918: 19–20 . Type material: BMNH 1922.8 .29.19, Holotype (seen). Material examined: SABAH – Crocker Range : BOR/ MOL 1292, Mahua , 1000 m alt., iii.2002 ; BOR/MOL 1307, Mahua , 1000 m alt., vii.2001 ; BOR/MOL 3608, Mahua , 1000 m alt., iv.2005 ; JJ 9721 , Mahua , iv.2002 . Diagnostic characteristics: The densely placed nodules over the surface and more angular periphery distinguish this species from E. paulbasintali and the others. Description: Shell ( Figs 6B , 10F, H ): large, rather thin, yellowish brown. Spire moderately elevated, outer whorls shouldered below the suture. Periphery slightly angular, more distinctly angular in juveniles. Above the periphery, shell with a silky lustre, very densely placed radial riblets, and cut by very densely arranged spiral grooves, forming nodules over the surface. Below the periphery, shell has fine, densely placed spiral grooves. Height up to 15.4 mm ; width up to 30.8 mm ; diameter of the first three whorls 1.3–1.5, 0.9–1.1, and 1.9–2.0 mm, respectively; number of whorls up to five and seven-eighths; height aperture up to 11.1 mm ; width aperture up to 15.8 mm . Genitalia ( Fig. 7A ): maximum length from genital opening to the end of dart-sac (before the visible gland tubules) up to 18 mm . The penis, dartsac, and vagina with thick muscular walls. P, V arranged near the GO then followed by BC and DS where there is much space in the atrium between the openings of P + V and BC + DS. Long BC about two-thirds of total DS length. Animal ( Figs 4A , 8I ): lighter black bands on either side of tentacles, which are divided by a white band that occupies the space between eye tentacles, and black bands faded towards the end of mantle and downward to just below the eye tentacles. Other parts of the animal are covered by regularly spaced oblique dotted belts. The mantle is almost uniformly reddish brown and followed by regularly spaced radial darker brown bands at the last half whorl. Distribution and habitat: Primary forest, 1200– 1800 m alt. Sabah : Crocker Range – Mahua and Mesilau ( Fig. 5B ). Remarks: The type materials were obtained from a natural history dealer in 1904 ( Gude, 1918 ). Type locality was in Borneo with unknown exact location.