<document id="4EB05AC857645999B62B87FFE9968903" ID-CLB-Dataset="6279" ID-DOI="10.5852/ejt.2021.751.1381" ID-GBIF-Dataset="4e3af683-d8a6-40de-991b-c4584d66b8cc" ID-ISSN="2118-9773" ID-Zenodo-Dep="4906007" ID-ZooBank="1D27EA41-2396-463A-B1B2-0CCF79C9F3D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623068023717" checkinUser="felipe" docAuthor="Zamani, Alireza, Chatzaki, Maria, Esyunin, Sergei L. &amp; Marusik, Yuri M." docDate="2021" docId="03F987850361EF7F1C617E7AFD026FF5" docLanguage="en" docName="ejt-751_68-114.pdf" docOrigin="European Journal of Taxonomy 751 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.751.1381" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Scotophaeus Simon 1893" docType="treatment" docVersion="6" lastPageNumber="98" masterDocId="FFC0FFFD037FEF611E347F73FFC16D41" masterDocTitle="One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations" masterLastPageNumber="114" masterPageNumber="68" pageNumber="98" updateTime="1699000119853" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D29F54AD0EE043A42860ABEB8C792E6E">One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations</mods:title>
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<mods:namePart id="B23BF347D6D595D9E8677ED09F1F039B">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="AAD9022A0DF91E632CF9BCD205AC5907">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland. Department of Molecular Biology and Genetics, Democritus University of Thrace, Dragana, 68100 Alexandroupolis, Greece. Perm State University, Bukireva Street 15, Perm, 614600, Russia. Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, 685000, Russia. Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa.</mods:affiliation>
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<mods:namePart id="38833B205FBA42420B19622267476786">Chatzaki, Maria</mods:namePart>
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<mods:namePart id="9E9561597FC4DBAB752976FE018F9DED">Marusik, Yuri M.</mods:namePart>
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Genus 
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<emphasis id="B924EA810361EF7F1C9C7E7BFC806C62" bold="true" box="[680,833,264,291]" italics="true" pageId="30" pageNumber="98">Scotophaeus</emphasis>
Simon, 1893
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<emphasis id="B924EA810361EF7F1E897E38FE856C24" bold="true" box="[189,324,331,357]" pageId="30" pageNumber="98">Comments</emphasis>
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The two new species described below are tentatively placed in 
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<emphasis id="B924EA810361EF7F1DF97E09FBA46CD5" box="[973,1125,378,404]" italics="true" pageId="30" pageNumber="98">Scotophaeus</emphasis>
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due to the absence of males and the general taxonomic uncertainty surrounding this genus. According to 
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, 
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<emphasis id="B924EA810361EF7F1E8A7EB1FE936C9D" box="[190,338,450,476]" italics="true" pageId="30" pageNumber="98">Scotophaeus</emphasis>
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is a “cumulative taxon for medium-sized robust gnaphosids”, a problem that has not been resolved; the genus requires a full revision (
<bibRefCitation id="EFC14B620361EF7F1CF27E95FC5A6F40" author="Murphy J." box="[710,923,486,513]" pageId="30" pageNumber="98" refId="ref17893" refString="Murphy J. 2007. Gnaphosid Genera of the World. British Arachnological Society St Neots, Cambridgeshire." type="book" year="2007">Murphy 2007: 51</bibRefCitation>
). However, the similarity of most of the somatic characters of the new species with 
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<emphasis id="B924EA810361EF7F1CFB7D79FCA76F65" box="[719,870,522,548]" italics="true" pageId="30" pageNumber="98">Scotophaeus</emphasis>
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(e.g., habitus, shape of maxillae, cheliceral dentition, eye configuration [as illustrated in 
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: fig. 374], presence of scopulae on metatarsi I and II [a diagnostic character of the genus, according to 
<bibRefCitation id="EFC14B620361EF7F1D5B7D21FBEE6F2D" author="Levy G." box="[879,1071,593,620]" pageId="30" pageNumber="98" pagination="427 - 452" refId="ref17599" refString="Levy G. 1999. Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology 45: 427 - 452." type="journal article" year="1999">Levy 1999: 437</bibRefCitation>
]) as well as the form of the genital organs, with a wide epigynal depression and multi-chambered receptacles, indicate that this genus is the best candidate for generic assignment.
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