Revision and cladistic analysis of the Southeast Asian leaf-dwelling spider genus Calapnita Simon (Araneae, Pholcidae) Author Bernhard A. Huber text Zootaxa 2017 4219 1 1 63 journal article 37319 10.5281/zenodo.273086 0d2332a5-1968-49d7-a570-f31d386eccb3 1175-5326 273086 0FA0F51A-3868-4F13-A93D-E34CA5A689F8 Calapnita bario sp. nov. Figs 120–121 , 134–141 Diagnosis . Distinguished from other species of vermiformis group by very long procursus ( Figs 134 , 288 ); from most species (except C. bariengi ) also by distal cheliceral apophyses not bipartite ( Fig. 136 ); from most (except C. dayak ) also by long and slender palpal tibia (tibia length/width>2.5; Fig. 286 ); from several species also by narrow and distinct epigynal ‘knob’ ( Fig. 141 ), and by long internal female genitalia with oval pore plates converging anteriorly and close together ( Fig. 140 ). FIGURES 120–127 . Live specimens, Calapnita vermiformis group. 120–121 . C. bario sp. nov. , ♂ and ♀ with egg-sac from Bario . 122–123 . C. magaseng sp. nov. , ♂ and ♀ with egg-sac from Mount Penrissen. 124–125 . C. saluang Huber , ♂ from Berastagi and ♀ with egg-sac from Phanom Bencha. 126–127 . C. vermiformis Simon , ♂♂ from Mount Isarog and Mount Banahaw. FIGURES 128–133 . Live specimens, Calapnita vermiformis group. 128 . C. loksado sp. nov. , ♂ from Loksado . 129–130 . C. bohol sp. nov. , ♂ and ♀ with egg-sac from near Loboc. 131 . C. dinagat sp. nov. , ♂ (with mite; arrow) from near Libjo. 132– 133 . C. nunezae sp. nov. , ♀ with egg-sac, and ♂ and ♀ at night from Katibawasan. FIGURES 134–141 . Calapnita bario sp. nov. (ZFMK, Ar 16022). 134–135 . Left procursus, prolateral and retrolateral views (arrow points at ventro-distal sclerite). 136–137 . Right male chelicera, frontal and lateral views. 138 . Left palpal trochanter, retrolateral view. 139 . Left palpal femur, prolateral view. 140 . Cleared female genitalia, dorsal view. 141 . Epigynum, ventral view (arrow points at ‘knob’). Scale lines: 0.5 mm (134–135), 0.3 mm (136–141). Etymology . The species name is derived from the type locality; noun in apposition. Material examined. Holotype . MALAYSIA-BORNEO: ♂ , ZFMK ( Ar 16021), Sarawak , Bario , forest along river N of town (3.765– 3.771°N , 115.444– 115.448°E ), 1170–1250 m a.s.l. , 29.vii.2014 ( B.A. Huber , S.B. Huber ). Other material . MALAYSIA-BORNEO: 1♂ 3♀ 4 juvs, ZFMK (Ar 16022), same data as holotype ; 1♀ 2 juvs in absolute ethanol, ZFMK (Bor 234), same data . Description. Male ( holotype ) MEASUREMENTS. Total body length 5.2, carapace width 0.75. Leg 1: 31.8 (7.8 + 0.4 + 7.2 + 13.6 + 2.8), tibia 2: 5.8, tibia 3: 3.6, tibia 4: 6.2; tibia 1 L/d: 103. Distance PME-PME 200 µm, diameter PME 110 µm, distance PME- ALE ~30 µm; no trace of AME. COLOR. Entire animal pale gray to whitish, legs slightly yellowish with brown patellae and tibia-metatarsus joints. BODY. Habitus as in Fig. 120 ; ocular area barely elevated, each triad on very low hump; carapace without median furrow; clypeus unmodified; sternum slightly wider than long (0.52/0.46), unmodified. CHELICERAE. As in Figs 136–137 , apophyses near lamellae not bipartite, with pair of lateral processes; without modified hairs; without stridulatory ridges. PALPS. In general similar to C. vermiformis (cf. figs 139 and 140 in Huber 2011 ); trochanter apophysis as in Fig. 138 ; femur as in Fig. 139 , ventral processes very widely spaced (distal process at 53% of femur length; cf. Fig. 285 ); tibia length/diameter 0.92/0.33; procursus as in Figs 134–135 , very long, with simple ventro-distal flat sclerite; bulb length 0.49; embolus length 1.16; embolus tip with transparent fringes. LEGS. Without spines and curved hairs; few vertical hairs; retrolateral trichobothrium on tibia 1 at 1.5%; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 pseudosegments barely visible in dissecting microscope. Male (variation). Tibia 1 in other male: 7.4. Female . In general similar to male; eye triads slightly closer together (distance PME-PME 180 µm). Tibia 1 in 3 females : 6.1, 6.1, 6.2. Epigynum as in Fig. 141 , weakly sclerotized triangular plate with small and slender anterior ‘knob’; internal genitalia as in Fig. 140 , very long, with pore plates close together and far from posterior margin. Distribution . Known from type locality in eastern Sarawak only ( Fig. 283 ).