Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species Author López-García, Margarita M. Author Deloya, Cuauhtémoc text The Coleopterists Bulletin 2019 2019-03-25 73 1 127 141 http://dx.doi.org/10.1649/0010-065x-73.1.127 journal article 10.1649/0010-065X-73.1.127 1938-4394 5333118 urn:lsid:zoobank.org:pub:A5E82F63-4AC3-41BD-B1D7-78430C22A742 Tomarus laticaudus López-Garc´ıa and Deloya, new species Zoobank.org/ urn:lsid:zoobank.org:act: AD2C9C9C-89DE-4DE3-94EE-FD4F50CBCC7E ( Figs. 2–5 ) Type Material. 56, 16♀ . Holotype labeled “Corazal/ CZ / Pn / iv.27.11 / AugBusck ” (16, USNM ) . Paratypes : Same label data as holotype ( 263♀ USNM ) . “Corazal/ CZ / Pn / iv.7.12 / A. Busck / light” (16 USNM ) . “ Diablo Hts. / CZ / Pn / 3.v.1971 / W.E. Bivin ” ( 1♀ USNM ) . “Arc-light/ Ancon / C.Z. / Pan. / 12.v.11 / A.H. Jennings ” (16 UNSM , 4♀ USNM ) . “ Arc-light globe/ Ancon / C.Z. / Pan. / Apr.1911 / Kraft” ( 2♀ UNSM , 2♀ USNM ) . “ Arc-light globe/ Ancon / C.Z. / Pan. / 27. Apr.1911 / E.A. Schwarz ” ( 2♀ USNM ) . “Paraiso / C.Z. / iv.24.11 / August Busck ” ( 1♀ UNSM ) . “ Panama / 27.xii.1912 / W. Robinson” ( 1♀ USNM ) . Figs. 2–6. Tomarus laticaudus : 2) Habitus; 3) Parameres, frontal view; 4) Parameres, lateral view; 5) Female pygidium, ventral view. Tomarus laevicollis : 6) Female pygidium, ventral view. Scale bars = 2 mm (Figs. 2, 5–6), 0.5 mm (Figs. 3–4). Description. Holotype . Habitus as in Fig. 2 . Length 22.2 mm ; width across humeri 12.0 mm. Color brown to black, venter slightly lighter. Head: Frons coarsely rugose. Frontoclypeal region with 2 triangular tubercles separated by about 7 tubercle diameters. Clypeus narrowed towards apex, base 3 times as wide as apex. Clypeal teeth triangular, separated by 1 tooth diameter. Mandibles with 2 apical, acute teeth and a rounded basal lobe. Interocular distance equal to 3.3 times eye width. Antennal club short, subequal in length to antennomeres 2–7. Pronotum: Surface smooth on disc, with minute punctures on lateral margins. Apical tubercle small and rounded. Fovea shallow, striate, and about half as wide as interocular distance. Margin of apical angles homogeneously rounded, not emarginate. Elytra: First interval punctate, punctures smaller than on other intervals. Pygidium: Surface rugose on basal fourth and lateral angles, disc with minute punctures. Slightly convex in lateral view. Apex regularly rounded. Legs: Protibia tridentate with an additional small basal tooth. Protarsus subcylindrical, not enlarged; inner claw entire. Metatibia not narrowed before apex, sides nearly parallel. Apex of metatibia crenulate, with 11 spinules. Parameres: With 2 pairs of dorsal lateral teeth; basal teeth long, wide, and upwardly arched, apical teeth short and joined to the basal ( Figs. 3–4 ). Variation. Male ( 4 paratypes ): Length 18.8–23.3 mm ; width across 9.9–12.1 mm . Female ( 16 paratypes ): Length 19.9–24.4; width across 10.0– 12.2 mm ; apex of pygidium dilated apically ( Fig. 5 ); apex of metatibia with 10–12 spinules. Etymology. The epithet laticaudus comes from the Latin words lati (large) and cauda (tail), which refer to the enlarged pygidium of the females. Distribution. Tomarus laticaudus is known from four localities in the former Canal Zone in Panama ( Fig. 23 ), which correspond to tropical moist forest areas at 10–80 m elevation in Panamá province . Temporal Distribution. April (14), May (6), December (1). Diagnosis. Tomarus laticaudus can be recognized by the nearly impunctate pronotum, small pronotal tubercle and fovea, and protibia with three teeth and an additional small basal tooth. It is similar to Tomarus laevicollis (Bates) , Tomarus colombianus López-Garc´ıa and Gasca-Álvarez, and Tomarus maracaiboensis López-Garc´ıa and Deloya, new species , but it can be differentiated by the strongly dilated pygidium of the females ( Fig. 5 ), which is homogeneously rounded in the other species ( e.g ., T. laevicollis ( Fig. 6 ), and the wide basal teeth of the parameres ( Figs. 3–4 ).