Resolving phylogenetic and classical nomenclature: A revision of Seraphsidae Jung, 1974 (Gastropoda: Neostromboidae) Author Maxwell, Stephen J. 0000-0002-4328-6171 College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. & Stephen. maxwell @ my. jcu. edu. au; https: // orcid. org / 0000 - 0002 - 4328 - 6171 tephen.maxwell@my.jcu.edu.au Author Rymer, Tasmin L. 0000-0002-9963-6345 College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. & Centre for Tropical Environmental and Sustainability Sciences, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. & Tamsin. rymer @ jcu. edu. au; https: // orcid. org / 0000 - 0002 - 9963 - 6345 amsin.rymer@jcu.edu.au Author Congdon, Bradley C. 0000-0002-8751-0892 College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. & Centre for Tropical Environmental and Sustainability Sciences, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. & brad. congdon @ jcu. edu. au; https: // orcid. org / 0000 - 0002 - 8751 - 0892 brad.congdon@jcu.edu.au text Zootaxa 2021 2021-06-22 4990 3 401 453 journal article 5563 10.11646/zootaxa.4990.3.1 6a6dd89d-6b74-47ed-ba8e-71b526355b69 1175-5326 5026777 70610FEE-4497-4804-817C-CEC2D66DDBFE Seraphs chilophorus ( Cossmann, 1889 ) Figure 4D 1889 Terebellum ( Seraphs ) chilophorum Cossmann , p. 94, pl. 3, figs. 5, 6. Jung 1974 , p. 22 . Caze et al. 2010 , p. 440 . = Seraphs ( Seraphs ) chilophorus Cossmann — Jung 1974 , p. 22, pl. 5, figs. 12–21, tfs. 15, 16. = Seraphs chilophorus Cossmann — Caze et al. 2010 , p. 440, figs. 15A–I. 2008 Seraphs volutatus Quaggioto & Mellini , p. 48, pl. V, fig. 29. Type material. Neotype — MNHN A25009 ; the neotype was selected by Caze et al. (2010) to bring taxonomic clarity, given the Cossmann (1889) type is missing. The latter specimen measures 29 mm in length, with a width of 8 mm , according to Jung (1974) . Type locality. Chaussy , France [Lutetian] . Diagnosis. The shell is moderately slender. The body whorl has a narrow, obsolete and longitudinal sulcus. The teleoconch has longitudinal sulci that are posteriorly reflected becoming axial, and terminating at the peristome. The aperture is smooth, and the labium has a salient narrow columella that is well developed and separated from the shell surface. The labrum basally runs parallel to the columella but gradually becomes confluent forming a posterior sinus, is marginated at the peristome, and sinuous adapically. The posterior sinus is decumbent becoming apically first sinistrally deflected, then reflected and recurved over the apex where it terminates dorsally. The columella is anteriorly recurved and extends further than the labrum. Distribution. EOCENE—Bartonian/Lutetian— France Les Boves, Paris Basin ( Jung 1974 ). Bartonian— France Caumont, Seine-et-Marne ( Caze et al. 2010 ). Italy Fontana del Cavaliere ( Eames 1952 ); Monte Postale, Ci- uppio, Valsugana and Croce Grand; San Giovanni, Ilarione ( Eames 1952 ; de Gregorio 1880 ); Sorne ( Eames 1952 ). Lutetian— France Chaussy, Val d’Oise ( Jung 1974 ; Caze et al. 2010 ); Saint-Lubin-de-la-Haye, Eure-et-Loir ( Caze et al. 2010 ); Fontenay-en-Vexin, Eure ( Caze et al. 2010 ); Vandancort, Oise ( Caze et al. 2010 ); Parnes, Oise ( Jung 1974 ; Caze et al. 2010 ); Fercourt, Oise ( Caze et al. 2010 ); Châteaurouge, Oise ( Caze et al. 2010 ); Saint-Félix, Oise ( Caze et al. 2010 ); Tiverval, Yvelines ( Caze et al. 2010 ); Grignon, Yvelines ( Jung 1974 ; Caze et al. 2010 ); Saulx- Marchais, Yvelines ( Jung 1974 ; Caze et al. 2010 ). Priabonian— Italy Cimome ( Eames 1952 ). Remarks. Seraphs chilophorus shared much of the Eocene with eight other species from Europe, all of which are distinguished from each other by subtle differences in form, sharing the typical lack of any distinctive sculpture, except for S. placiferus with distinctive axial growth cords. This morphology of S. chilophorus is similar to that of S. peterjungi and only distinguishable from that species by its residual colour.