The land snail genus Macrochlamys Gray, 1847 from Thailand, with descriptions of five new species (Pulmonata: Ariophantidae) Author Pholyotha, Arthit Author Sutcharit, Chirasak Author Panha, Somsak text Raffles Bulletin of Zoology 2018 2018-12-03 66 763 781 journal article 10.5281/zenodo.5460634 2345-7600 5460634 1E7214C5-24E8-4860-BF13-203FD93A2C19 Macrochlamys brunnea Möllendorff, 1902 ( Fig. 8A ) Macrochlamys brunnea Möllendorff, 1902b: 155 ( type locality: Bangkok , Muok Lek, Hinlap) – Hemmen & Hemmen, 2001: 44 . Material examined. Holotype ( SMF 227090 /1), Bangkok , Siam [ Bangkok , Thailand ] . Paratypes : one shell ( SMF 227091 /1), Bangkok , Thailand ; four shells ( SMF 227171 /4) , Table 1. Comparison of shell and genital system of Macrochlamys species in Thailand.

Characters	M. aurantia new species	M. coleus new species	M. caverna new species	M. lemma new species	M. tanymentula new species	M. brunnea	M. kelantanensis
Shell width (mm) ranges, means ± SD and no. of specimens	11.7–14.0 12.8 ± 0.98 (n=6)	10.2–13.0 11.6 ± 0.99 (n=6)	13.2–17.2 14.3 ± 1.09 (n=11)	13.1–17.0 14.7 ± 1.54 (n=6)	9.6–11.1 10.6 ± 0.58 (n=5)	9.0*	16.7–24.0 20.0 ± 2.56 (n=9)
Shell height (mm) ranges, means ± SD and no. of specimens	6.5–8.2 7.5 ± 0.66 (n=6)	5.7–6.6 6.0 ± 0.35 (n=6)	7.3–8.6 7.6 ± 0.51 (n=11)	6.5–7.6 7.1 ± 0.40 (n=6)	4.9–5.5 5.2 ± 0.27 (n=5)	5.0*	9.0–13.0 11.4 ± 1.21 (n=9)
Shape of suture	Deep channel	Shallow channel	Impressed	Impressed	Impressed	Impressed	Impressed
Umbilicus	Narrowly open	Narrowly open	Widely open	Narrowly open	Narrowly open	Narrowly open	Narrowly open
Penial caecum	Present	Present	Present	Present	Absent	–	Absent
Inner wall of penis sculpture (proximal / distal)	Both with small wrinkles	Small wrinkled / small longitudinal and wrinkled folds	Trapezoid / lateral folds	Trapezoid / lateral folds	Both with weak wrinkles	–	Both with weak wrinkles
Epiphallus	Short	Short	Short	Rather long	Very long	–	Slightly long
Epiphallic caecum	One circle	One circle	One circle	One circle	One circle	–	Two circles
Tissue casing	Absent	Present	Absent	Present	Absent	–	Absent
Vagina length	Long	Slightly long	Long	Slightly long	Short	–	Short
Position of thicken tissue on free oviduct	Base	Base	One-fourth from base	Near middle	Base	–	Base
Location of dart apparatus	Proximal end of vagina	Proximal end of vagina	Proximal end of vagina	Near proximal end of vagina	Proximal end of vagina	–	Proximal end of vagina

*Shell measurement taken from the original description of Möllendorff (1902b) . Muok Lek, Siam [Muak Lek District, Saraburi Province , Thailand ]. Remarks. Macrochlamys brunnea differs from its congeners in Thailand by its small size and globose shell ( Table 1 ). Other distinguishing characters of this species are the depressed globose, shiny dark brown shell with a smooth surface and rounded periphery. The shell has a low conical spire, which is rather convex, with a slightly raised apex, and shallow suture. Its aperture is ovate-lunate in shape, opens obliquely and has a simple peristome. The columellar margin is gradually reflected near the narrowly open and very small umbilicus ( Fig. 8A ). The generic placement of this species is tentative and requires verification since anatomical data is thus far unavailable. It is herein provisionally retained in the Macrochlamys as originally proposed although this species can be assigned to either Sarika or Macrochlamys if based solely on shell morphology. Unfortunately, attempts to obtain topotypic specimens from Bangkok and accessible limestone outcrops in the Saraburi Province were unsuccessful and no specimens identifiable as M. brunnea could be found. This could be due to habitat loss and population decline as many limestone outcrops in this region are being quarried. Its current status is unclear.