New genera and new species of earthworms (Oligochaeta: Octochaetidae) from Western Ghats, South India Author Julka, J. M. Author Blanchart, Eric Author Chapuis-Lardy, Lydie text Zootaxa 2004 486 1 27 journal article 10.5281/zenodo.157661 65483b24-e197-4508-9c49-50fb2a6ec291 1175­5326 157661 48073126-2C08-4E8A-8150-00C9DB50EBD6 Herbettodrilus gen. nov. Diagnosis. Prostomium prolobic. Setae lumbricine. Male pores paired, in seminal grooves, on 18; prostatic pores paired, at ends of seminal grooves, on 17 and 19. Oesophagus with a single gizzard, in 5 and one pair of discrete, extramural calciferous glands, each gland bilobed, one lobe in 15 and the other in 16, both discharging into gut through a common duct in 15 close to attachment of septum 14/15; intestinal caeca and supra­intestinal glands absent; typhlosole simple, lamelliform. Penial setae present. Micromeronephridia astomate, enteronephric paired tufts in 3, few scattered on body wall in 4 and posteriad segments; paired, stomate, enteronephric megameronephridia in 15 and posteriad segments, discharging into rectum through paired excretory canals. Herbettodrilus belongs to a group of Oriental octochaetid genera with a single oesophageal gizzard, discrete extramural calciferous glands and stomate enteronephric megameronephridia. It is distinguished from other members of the group, Hoplochaetella Michaelsen, 1900 and Karmiella Julka, 1983 by the characteristics as given in Table 3 . TABLE 3. Comparison of Herbettodrilus with related genera.

Characteristics	Herbettodrilus	Karmiella	Hoplochaetella
Prostomium	prolobic	tanylobic	epilobic
Setae	lumbricine	lumbricine	perichaetine
Male terminalia	male pores one pair, in seminal grooves, on 18; prostatic pores two pairs, at ends of seminal grooves, on 17 and 19	male pores one pair, in seminal grooves, on 18; prostatic pores two pairs, at ends of seminal grooves, on 17 and 19	combined male and pros­ tatic pores two pairs, on 17 or 17/18 and 19 or 18/19; rarely one pair, on 17; sem­ inal grooves absent
Gizzard	in 5	in 5	in 6 or 7
Calciferous glands	one pair; each gland bilobed, one lobe in 15 and the other in 16, both dis­ charging into oesophagus through a common stalk close to attachment of sep­ tum 14/15	one pair; each gland bilobed, one lobe in 15 and the other in 16, both dis­ charging into oesophagus through a common stalk close to attachment of sep­ tum 15/16	four pairs, in 10–13
Paired stomate enter­ onephric megamero­ nephridia	in 15 and posteriad seg­ ments, each with one fun­ nel	from about middle of body to posteriormost segment, each with one funnel	in 12–14 and posteriad segments, each with one functional and several ves­ tigial secondary funnels

Type species. Herbettodrilus bahli sp. nov. Description. As for the only species, bahli . Etymology. Herbettodrilus , gender masculine; derived from the locality ‘Herbettu’, and the Greek word ‘drilos’, meaning the worm.