Revision of Ephydrini Zetterstedt (Diptera: Ephydridae) from the Americas south of the United States Author Mathis, Wayne N. Author Marinoni, Luciane text Zootaxa 2016 4116 1 1 110 journal article 38982 10.11646/zootaxa.4116.1.1 c41804b4-7a66-4a00-afc1-c7a31871329f 1175-5326 257322 22D15539-E49E-4D6C-BFCF-D4DBC72BA640 5. Paracoenia (Paracoenia) wirthi Mathis Figs. 27–34 , Map 5 Paracoenia (Paracoenia) wirthi Mathis 1975 : 78 [ United States . California. Inyo: Tecopa Hot Springs ( 1.6 km N); HT ♂, CAS (12033)].— Mathis 1980 : 9 –11 [revision].— Mathis and Zatwarnicki 1995 : 251 [world catalog]. Diagnosis. This species is distinguished from congeners, especially P. bisetosa , by the following combination of characters: Medium-sized shore flies, body length 3.12–3.65 mm ; subshiny dorsally, bluish green to green metallic luster. Head ( Figs. 27–28 ): Mesofrons greenish blue, shiny; fronto-orbital areas dark brown, microtomentose to subshiny. Face densely microtomentose, grayish tank, dorsum of facial hump slightly darker than face, grayer; paravertical setae shorter than their distance apart; gena-to-eye ratio 0.30; eye ratio 0.64, height-to-length ratio 0.91; eye-width-to-face-length ratio 0.44. Thorax: Mesonotum ( Fig. 29 ) with anterior area slightly microtomentose, becoming subshiny to shiny posteriorly; acrostichal setulae in 3–4 rows anteriorly, becoming irregular, 5–6 rows posteriorly. Mesopleuron with more subdued coloration than mesonotum, mostly olivaceous gray. Wing nearly transparent to faintly brown. Abdomen: Concolorous with mesonotum; digitiform lateral process of male tergite 5 elongate, acutely pointed apically; male sternite 5 ( Fig. 34 ) broadly U-shaped with posteriorly extended arms oriented posteriorly, acutely pointed, anterior margin somewhat truncate, posterior margin moderately deeply emarginate. Male terminalia ( Figs. 30–32 ): Surstylus in posterior view with base robustly developed, width 3X apical width, thereafter abruptly narrowed with apical extension narrowly digitiform; medial triangular process less than half length of surstylar arm, not cleft apicomedially; gonite more or less triangular, sider basally, apical half tapered, apex digitiform, apex rounded. Female ventral receptacle as in Fig. 33 with large, mushroom-like operculum. FIGURES 27–29. Paracoenia wirthi Mathis. (27) head, anterior view; (28) same, lateral view; (29) mesonotum, dorsal view (from Mathis 1980). FIGURES 30–34. Structures of male and female terminalia of Paracoenia wirthi Mathis. (30) epandrium, cerci and surstyli, posterior view; (31) epandrium, cerci, surstyli and gonite, lateral view; (32) ventral apex of male tergite 5, ventral view; (33) female ventral receptacle, lateral view; (34) male sternite 5, ventral view (from Mathis 1980). MAP 5. Distribution map for Paracoenia wirthi Mathis. Type material. The holotype male is labeled “ USA . CALIF . Inyo Co 1 mi N Tecopa Hot Spg 24 June 1974 [,] Wayne N. Mathis/ HOLOTYPE Paracoenia wirthi Mathis [red; 12033].” Allotype and 21 paratypes ( 7♂ , 14♀; CAS , USNM ) bear the same label data as the holotype . Other paratypes as listed in Mathis (1975:78) . The holotype male is double mounted (minute nadel), is in excellent condition, and is deposited in the CAS (12033). Type locality. United States . California. Inyo: Tecopa Hot Springs ( 1.6 km N; 35°52.7'N , 116°13.9'W ); HT ♂, CAS (12033). Other specimens examined. MEXICO . Baja California: Guadalupe Canyon ( 32°09.3'N , 115°47.4'W ), 19 May 1957 , F. X. Williams ( 1♂ , 3♀; CAS ). Distrito Federal: Mixquic, Chalco Lake ( 9 km NW; 19°15.5'N , 98°58.6'W ), 1 Aug 1965 , K. R. Valley ( 4♂ , 16♀; CAS , USNM ). Distribution (Map 5). Nearctic: United States (California). Neotropical: Mexico (Baja California, Distrito Federal). Remarks. This species is similar to P. bi s pi no s a but is slightly smaller in body length and less setulose. The ventral margin of the male sternite five is broadly U-shaped, and the connecting bridge is wide; the male tergite five is produced into an acutely pointed process; and the surstylus is more robustly developed basally, and the distal surstylar extensions are narrowly digitiform; the gonite is narrowed toward the apex, and the apex is digitiform, apically rounded.