Review of the Pacific leafhopper genus Hecaloidella (Hemiptera: Cicadellidae Deltocephalinae: Selenocephalini) with description of sixteen new species Author Xu, Deliang 0000-0003-0106-5570 Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A & F University, Yangling, Shaanxi 712100, China & 18710305867 @ 163. com; https: // orcid. org / 0000 - 0003 - 0106 - 5570 Author Webb, Michael D. 0000-0002-1312-6142 Department of Life Sciences, Natural History Museum, Cromwell Road, London, ME 15 6 QX, UK & mdwebb 04 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1312 - 6142 mdwebb04@gmail.com Author Shang, Suqin 0000-0001-9376-0735 Biological Engineering Laboratory of Crop Diseases and Pests of Gansu Province, College of Plant Protection, Gansu Agricultural University, Lanzhou, Gansu 730070, China & shangsq @ gsau. edu. cn; https: // orcid. org / 0000 - 0001 - 9376 - 0735 shangsq@gsau.edu.cn Author Zhang, Yalin 0000-0002-1204-9181 Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A & F University, Yangling, Shaanxi 712100, China & yalinzh @ nwsuaf. edu. cn; https: // orcid. org / 0000 - 0002 - 1204 - 9181 yalinzh@nwsuaf.edu.cn text Zootaxa 2021 2021-03-22 4948 3 381 403 journal article 7470 10.11646/zootaxa.4948.3.4 6bfd846c-0304-4330-8941-95c406630230 1175-5326 4629100 8E0BF97A-D9E1-4F2E-BB63-82A6ED47E5DC H. spinosa Zhang & Webb sp. nov. Figs. 1M , 6P–U Description. Body length (including tegmina), male: 5.2mm ; female: 5.2mm . Vertex pale yellow with a pair of short reddish brown longitudinal bars joined basally. Face brownish yellow. Pronotum brownish yellow, paratype with darker brown irroration. Forewing veins brown, more heavily marked with brown on some veins including a spot on apex of claval veins, on veins of fifth apical cell and on apical cells and on wing apex. Male genitalia with subgenital plate strongly constricted sub-basally with a long digitate apex with moderately long fine setae. Connective with arms long and appressed; stem short with a pair of apical spines. Aedeagus firmly attached to connective; preatrium with a long ventral process, distally bifurcate; shaft elongate with a pair of widely divergent apical processes, gonopore apical. Female with pregenital sternite strongly produced medially ( Fig. 6U ). Material examined. Holotype : ♂ , Brunei , Ulu Temburong Ridge, ii-iii,1982, M.C. Day , NHMUK 010591778 ( BMNH ) . Paratype : 1♀ , Brunei , Ulu Temburong , 300m ., ii-iii.1982, M.C. Day ( BMNH ) . Etymology. This species is named after the pair of spines on the apex of connective stem. Remarks. This species can be distinguished from all other Hecaloidella species by the unique shape of the connective and aedeagus (see description) and in the female by the greatly prolonged pregenital sternite posteromedially.