Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae) Author Selnekovič, Dávid D930D757-D929-4248-A29C-A857E0505345 Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia. david.selnekovic@uniba.sk Author Jäch, Manfred A. D749707A-8823-4110-8D51-BF5AF4E2820F Naturhistorisches Museum Wien, Burgring 7, A- 1010 Wien, Austria. manfred.jaech@nhm-wien.ac.at Author Kodada, Ján 6E88BFBB-8769-44EC-8285-29E357CEC064 Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia. jan.kodada@uniba.sk text European Journal of Taxonomy 2024 2024-09-11 957 1 229 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2651/12247 journal article 10.5852/ejt.2024.957.2651 2118-9773 13773385 9957F2DF-3F31-4B58-84BC-2F8E0F45C6E3 Elmomorphus umphangicus Kodada, Selnekovič & Jäch sp. nov. urn:lsid:zoobank.org:act: 309D3D42-6B56-4D27-B75A-7B8F80FD8B03 Figs 44–45 , 111C Differential diagnosis Elmomorphus umphangicus sp. nov. ( Fig. 44 ) is characterised by having a plastron on the cranial surface (except for a small triangular area on the vertex) and on the lateral sides of the pronotum and elytra. Elytral punctures are scattered over the entire surface and longitudinal striae are absent. Such a combination of characters is also present in E. bispinosus sp. nov. , E. longitarsis sp. nov. , E. parvulus sp. nov. , E. sausai sp. nov. , and E. yunnanensis sp. nov. The elytral plastron bands are slenderer than in E. sausai . Elytra are widest behind the middle in E. umphangicus , while in the rest of the species, they are widest around the middle. Elmomorphus umphangicus is smaller (TL: ♂♂ 2.64–2.90 mm (2.76±0.10, n=5), ♀♀ 2.46– 3.12 mm (2.83 ±0.30, n=6)) than E. bispinosus (TL: ♀♀ 3.38–3.54 mm (n=3)), E. longitarsis sp. nov. (TL: ♂♂ 2.93–3.09 mm (n =3)), E. sausai (TL: ♂♂ 3.07–3.15 mm (3.10±0.03, n=4)), or E. yunnanensis (TL: ♂♂ 3.29–3.67 mm (3.37 ±0.15, n=4)) and slightly larger than E. parvulus (TL: ♂♂ 2.41–2.62 mm (2.54±0.07, n=6), ♀♀ 2.60–2.81 mm (2.71± 0.07, n=6)). Pronotal and elytral surface with distinct bronze lustre ( Fig. 44 ), not present in E. yunnanensis ( Fig. 50 ). Aedeagus as in Fig. 45 . Etymology The epithet is a Latin adjective in the nominative singular referring to the type locality of the species. Type material Holotype THAILAND – Tak Province • ♂ ; “THAI, 26.IV.-6.V.1991 UMPHANG 500 m 16°04′N 98°53′E Vít Kubáň leg.”; NMW . Paratypes THAILAND – Tak Province • 2 ♂♂ , 6 ♀♀ ; same collection data as for holotype; CKB , NMW • 3 ♀♀ ; with similar labels, but “David Král lgt.”; NMW . Fig. 44. Elmomorphus umphangicus Kodada, Selnekovič & Jäch sp. nov. , paratype, male from type locality (CKB), TL: 2.80 mm. Fig. 45. Elmomorphus umphangicus Kodada, Selnekovič & Jäch sp. nov. , aedeagus. A . Holotype (NMW), lateral aspect. B . Holotype (NMW), ventral aspect. C . Paratype from type locality (CKB), lateral aspect. Scale bar: 0.1 mm. Remarks A series of 67 specimens (NMW) labelled “ Thailand 2003 Prov. Suratthani [ Surat Thani ] Khao Sok N.P. 15.1. leg.: Horst Forster”, closely resembles the type specimens of E. umphangicus sp. nov. , but they differ in the slightly more convex elytra and the slightly narrower elytral plastron bands. Although we could not detect any apparent differences in the male genitalia, we are not including these specimens from Khao Sok N.P. ( Surat Thani Province ) in the type series or as “additional material examined” of E. umphangicus ; molecular data will be necessary to reveal the identity of these specimens. Type locality Thailand , Tak Province , Umphang, 16°4′ N , 98°53′ E . Description Measurements (mm): TL: ♂♂ 2.64–2.90 (2.76± 0.10, n= 5), ♀♀ 2.46–3.12 (2.83 ± 0.30, n =6); PL: ♂♂ 0.61–0.66 (0.63± 0.02, n =5), ♀♀ 0.53–0.71 (0.63± 0.08, n =6); PW: ♂♂ 1.06–1.17 (1.11 ±0.04, n=5), ♀♀ 0.96–1.26 (1.11 ±0.13, n=6); EL: ♂♂ 2.03–2.24 (2.13± 0.07, n=5), ♀♀ 1.92– 2.42 (2.20 ± 0.22, n =6); EW: ♂♂ 1.30–1.43 (1.35± 0.05, n=5), ♀♀ 1.17–1.51 (1.36 ± 0.14, n =6); PhL: 0.53–0.64 (0.60± 0.05, n =5); PrL: 0.31–0.34 (0.32 ±0.01, n =5). Body elongate oval, widest behind midlength of elytra, moderately convex dorsally ( Fig. 44 ). Colouration black; mouthparts, antennae, trochanters, and tarsi reddish brown, remaining parts of legs dark brown. Dorsal pubescence consists of thin short decumbent setae arising from small round punctures. Dorsal plastron on cranium except for triangular area on vertex, on lateral portions of pronotum and elytra. Plastron present on ventral surface, except for prosternal process, median part of metaventrite, and middle of ventrites 1–2. Cranium with round setiferous punctures smaller than eye facet and separated by 0.5–1.0 × puncture diameter. Plastron on dorsal cranial surface, except for triangular area on vertex. Labrum transverse with straight anterior margin; exposed portion microreticulate with small setiferous punctures; setae concentrated mainly along anterior margin. Anterior margin of clypeus straight. Eyes large, oval, moderately protruding, ID: ♂♂ 0.37–0.42 mm (0.40± 0.02, n =5), ♀♀ 0.34–0.45 mm (0.41± 0.04, n= 6). Antennae 10-segmented, densely setose. Pronotum transverse, weakly convex, widest at base, PW/PL: ♂♂ 1.74–1.77 (1.76± 0.01, n=5), ♀♀ 1.71– 1.83 (1.77± 0.05,n =6). Plastron forming medially constricted band along lateral pronotal side. Remaining pronotal surface smooth, with round setiferous punctures slightly larger than those on head and subequal elytral punctures. Anterior pronotal angles protruding and deflexed; lateral pronotal sides convergent, straight, or weakly rounded. Prosternal process with lateral and posterior edges rounded; lateral portions weakly raised, without clusters of long setae; median keel flat. Scutellum longer than wide, smooth, with several setiferous punctures. Metaventral process with lateral margins slightly raised; metaventral disc flat, without clusters of long setae; posteriorly widened median portion smooth, without plastron. Elytra ovate, moderately convex, widest behind midlength, EL/EW: ♂♂ 1.56–1.63 (1.58 ±0.03, n=5), ♀♀ 1.58–1.64 (1.62 ± 0.02, n= 6). Plastron in two posteriorly expanded lateral bands, each covering about half of elytron width (along elytral midlength); remaining elytral surface weakly microreticulate, with round setiferous punctures. Tibiae straight; protibia as long as protarsus; PrTL/PL: ♂♂ 1.17–1.28 (1.21 ± 0.05, n=5), ♀♀ 1.17–1.34 (1.25± 0.07, n= 6). Terminal protarsomere as long as all preceding segments combined. Male foreclaws narrow, strongly curved, similar to those of female. Ventrites covered with plastron, except for middle of ventrites 1–2. Admedian keels on ventrite 1 inapparent. Ventrite 5 evenly convex in both sexes; apex in males slightly excised, in females rounded with short longitudinal keel. Aedeagus ( Fig. 45 ): phallobase rather short, PhL/PrL: 1.69–2.00 (1.85– 0.14, n=5); parameres narrowed and rounded apically (lateral view), weakly curved; penis narrowly rounded at apex; sclerotised fibula slender. Secondary sexual dimorphism Apex of male ventrite 5 shallowly excised, in female rounded, with short longitudinal keel. Distribution Thailand ( Fig. 111C ).