<document id="63824C126A047EF13FC2FBFD26FDC85E" ID-DOI="10.5281/zenodo.5353792" ID-ISSN="1942-1354" ID-Zenodo-Dep="5353792" ID-ZooBank="88D45C5E-0E49-4F02-87A4-C4FA55A83B2F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630212659467" checkinUser="felipe" docAuthor="Freese, Edwin L., Veal, Doug A. &amp; Lago, Paul K." docDate="2020" docId="28608792FFB7364AFF76FD08BAF9FB03" docLanguage="en" docName="0787.pdf" docOrigin="Insecta Mundi 2020 (787)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Tomarus relictus" docType="treatment" docVersion="3" lastPageNumber="48" masterDocId="D459FFEAFF86367BFFB6FFEDBB37FF84" masterDocTitle="The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist" masterLastPageNumber="83" masterPageNumber="1" pageNumber="48" updateTime="1699188000474" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="8341BDF4781AAF732CDF7066120C009A">The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist</mods:title>
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<mods:namePart id="F6CA849C630440DC16D0A44CDADD8DE7">Freese, Edwin L.</mods:namePart>
<mods:affiliation id="A34883FC258E21AC74E2EE76035D80F5">33493 “ S ” Avenue Adel, Iowa 50003</mods:affiliation>
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<taxonomicName id="67C94D07FFB7364AFF76FD08B973FD78" authority="(Say, 1825)" baseAuthorityName="Say" baseAuthorityYear="1825" box="[192,580,740,764]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="48" phylum="Arthropoda" rank="species" species="relictus">
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<emphasis id="92BDEA96FFB7364AFF76FD08BA9EFD78" bold="true" box="[192,425,740,764]" italics="true" pageId="49" pageNumber="48">Tomarus relictus</emphasis>
(Say, 1825)
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<paragraph id="A0763684FFB7364AFF76FCFCB91CFCC2" blockId="49.[192,1440,785,838]" pageId="49" pageNumber="48">
<bibRefCitation id="C4584B75FFB7364AFF76FCFCBA4AFCAD" author="Wickham, H. F." box="[192,381,785,809]" pageId="49" pageNumber="48" pagination="1 - 40" refId="ref34735" refString="Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40." type="journal article" year="1911">Wickham 1911</bibRefCitation>
; 
<bibRefCitation id="C4584B75FFB7364AFE3AFCFCB914FCAD" author="Hardy, A." box="[396,547,785,809]" pageId="49" pageNumber="48" pagination="1 - 57" refId="ref31814" refString="Hardy, A. 1991. A catalog of Coleoptera of America north of Mexico. Family: Scarabaeidae. Subfamilies Rutelinae and Dynastinae. Volume 34, part 2. United States Department of Agriculture Handbook 529: 1 - 57." type="journal article" year="1991">Hardy 1991</bibRefCitation>
; 
<bibRefCitation id="C4584B75FFB7364AFD84FCFCB887FCAD" author="Worthington, R. J. &amp; K. J. Larsen" box="[562,944,785,809]" pageId="49" pageNumber="48" pagination="77 - 90" refId="ref34793" refString="Worthington, R. J., and K. J. Larsen. 2010. An annotated checklist of scarab beetles (Coleoptera: Scarabaeidae) from northeastern Iowa. The Great Lakes Entomologist 43 (1 - 4): 77 - 90." type="journal article" year="2010">Worthington and Larsen 2010</bibRefCitation>
; 
<bibRefCitation id="C4584B75FFB7364AFC09FCFCBFC7FCAD" author="Ratcliffe, B. C. &amp; R. D. Cave" box="[959,1264,785,809]" pageId="49" pageNumber="48" pagination="1 - 298" refId="ref33638" refString="Ratcliffe, B. C., and R. D. Cave. 2017. The dynastine scarab beetles of the United States and Canada (Coleoptera: Scarabaeidae: Dynastinae. Bulletin of the University of Nebraska State Museum 30: 1 - 298." type="journal article" year="2017">Ratcliffe and Cave 2017</bibRefCitation>
; ISIC; IWUC; CCMV; DAVC; ELFC; PKLC.
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<paragraph id="A0763684FFB7364AFF76FCBBB95FFCEA" blockId="49.[192,616,854,878]" box="[192,616,854,878]" pageId="49" pageNumber="48">Collection dates: April–November.</paragraph>
<paragraph id="A0763684FFB7364AFF76FC90B995FC11" blockId="49.[192,674,893,917]" box="[192,674,893,917]" pageId="49" pageNumber="48">Collected at light and from horse dung.</paragraph>
<paragraph id="A0763684FFB7364AFF76FC48B8A9FB97" blockId="49.[192,1440,932,1043]" pageId="49" pageNumber="48">
County records (Figure 176): Appanoose, Black Hawk, Boone, Bremer, Carroll, Cerro Gordo, Cherokee, Dallas, Davis, Decatur, 
<collectingRegion id="620DF866FFB7364AFD93FC2CB9A9FC5D" box="[549,670,961,985]" country="United States of America" name="Delaware" pageId="49" pageNumber="48">Delaware</collectingRegion>
, Des Moines, Dickinson, Fremont, Henry, Jefferson, Johnson, Keokuk, Linn, Louisa, Lucas, Marion, Monona, Monroe, Muscatine, Osceola, Page, 
<collectingRegion id="620DF866FFB7364AFB52FC33BE68FC72" box="[1252,1375,990,1014]" country="United Kingdom" name="Plymouth" pageId="49" pageNumber="48">Plymouth</collectingRegion>
, Pottawattamie, Scott, Story, Van Buren, Wapello, Winneshiek.
</paragraph>
<paragraph id="A0763684FFB7364AFF76FBBFB979FBEE" blockId="49.[192,590,1106,1159]" box="[192,590,1106,1130]" pageId="49" pageNumber="48">
Tribe 
<taxonomicName id="67C94D07FFB7364AFEBCFBBFB979FBEE" authority="Mulsant, 1842" authorityName="Mulsant" authorityYear="1842" box="[266,590,1106,1130]" class="Insecta" family="Dynastidae" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="48" phylum="Arthropoda" rank="tribe" tribe="Oryctini">ORYCTINI Mulsant, 1842</taxonomicName>
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<taxonomicName id="67C94D07FFB7364AFF76FB82BAF9FB03" authority="Hope, 1837" authorityName="Hope" authorityYear="1837" box="[192,462,1135,1159]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="92BDEA96FFB7364AFF76FB82BA0BFB03" box="[192,316,1135,1159]" italics="true" pageId="49" pageNumber="48">Xyloryctes</emphasis>
Hope, 1837
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