The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist Author Freese, Edwin L. 33493 “ S ” Avenue Adel, Iowa 50003 Author Veal, Doug A. 168 Thompson Drive SE Cedar Rapids, Iowa 52403 Author Lago, Paul K. Biology Department, University of Mississippi University, Mississippi 38677 text Insecta Mundi 2020 2020-08-28 2020 787 1 83 journal article 10.5281/zenodo.5353792 1942-1354 5353792 88D45C5E-0E49-4F02-87A4-C4FA55A83B2F Geotrupes opacus Haldeman, 1853 Putnam 1876 ; Wickham 1911 ; Howden 1955a ; ISIC; IWUC; ELFC; DAVC; MJPC. Collection dates: April, September–October. Specimens were collected from dung and on log. Specimen label data: Lacey Keosauqua State Park, Van Buren Co., Iowa , 14 October 2005 , on log, ELF (ELFC). County records ( Figure 13 ): Black Hawk, Cedar, Dallas, Henry, Johnson, Linn, Louisa, Pottawattamie, Story, Van Buren, Woodbury.