A revision of Geonoma (Arecaceae) Author Henderson, Andrew text Phytotaxa 2011 2011-02-18 17 1 271 journal article 24911 10.11646/phytotaxa.17.1 65962ba7-eec1-40e7-aed6-cef94e99ca39 1179-3163 3538362 40. Geonoma multisecta (Burret) Burret (1940a: 24) . Taenianthera multisecta Burret (1930c: 13) . Type : COLOMBIA . Caquetá : Sucre, 10 July 1926 , G . Woronow & J . Juzepczuk 5862 ( holotype LE !). Geonoma polyandra Skov (1994: 39) . Type : ECUADOR . Napo : Añangu, 0°32’N , 76°23’W , 300 m , 19 June 1985 , H. Balslev, A. Barfod, & F. Skov 60536 ( holotype AAU!, isotypes COL n.v. , K!, NY!, QCA n.v. ), s ynon. nov. Plants 2.7(1.5–3.5) m tall; stems 0.8(0.2–2.3) m tall, in diameter no data, solitary; internodes length no data, not scaly. Leaves 11(9–16) per stem, irregularly pinnate, not plicate, bases of blades running diagonally into the rachis; sheaths 22.0(11.0–38.0) cm long; petioles 41.2(13.0–77.0) cm long, drying green or yellowish; rachis 74.6(55.5–94.0) cm long, 4.3(2.9–8.0) mm in diameter; veins raised and rectangular in cross-section adaxially; pinnae13(3–18) per side of rachis; basal pinna 29.0(21.0–54.0) cm long, 2.5(0.2–18.5) cm wide, forming an angle of 56(20–74)° with the rachis; apical pinna 20.6(15.5–29.5) cm long, 5.7(2.0–17.0) cm wide, forming an angle of 32(22–42)° with the rachis. Inflorescences unbranched; prophylls and peduncular bracts ribbed with elongate, unbranched fibers, both bracts tubular, narrow, elongate, closely sheathing the peduncle, more or less persistent; prophylls 29.1(19.2–40.0) cm long, not short and asymmetrically apiculate, the surfaces not ridged, without unequally wide ridges; peduncular bracts 29.3(25.0–39.5) cm long, well- developed, inserted 0.8(0.2–2.5) cm above the prophyll; peduncles 60.8(31.0–99.5) cm long, 3.7(2.3–5.1) mm in diameter; rachillae 1, 23.5(14.0–36.5) cm long, 7.4(5.4–9.4) mm in diameter, the surfaces without spiky, fibrous projections or ridges, drying brown or yellow-brown, without short, transverse ridges, not filiform and not narrowed between the flower pits; flower pits spirally arranged, glabrous internally; proximal lips with a central notch before anthesis, often the two sides of the notch overlapping, not recurved after anthesis, not hood-shaped; proximal and distal lips drying the same color as the rachillae, not joined to form a raised cupule, the proximal lip margins overlapping the distal lip margins; distal lips well-developed; staminate and pistillate petals not emergent, not valvate throughout; staminate flowers deciduous after anthesis; stamens more than 6; thecae diverging or not diverging at anthesis, inserted onto poorly to well-developed, non-split, jointed connectives, connectives when well-developed alternately long and short; anthers short at anthesis, remaining straight and parallel; non-fertilized pistillate flowers deciduous after anthesis; staminodial tubes lobed at the apex, the lobes spreading at anthesis, acuminate, those of non-fertilized pistillate flowers not projecting and persistent after anthesis; fruits 11.2(9.6–13.2) mm long, 7.4(6.2–7.9) mm in diameter, the bases without a prominent stipe, the apices not conical, the surfaces not splitting at maturity, without fibers emerging, not bumpy, not apiculate; locular epidermis with operculum, smooth, without pores. Distribution and habitat:— From 1°47’N– 2°17’S and 74°53’– 77°45’W in the western Amazon region of Colombia and Ecuador at 338(200–1150) m elevation in lowland or) montane rainforest ( Fig. 28 ). FIGURE 28 . Distribution maps of Geonoma monospatha , G . mooreana , G . multisecta , and G . occidentalis . Taxonomic notes:— Skov (1994) , in describing Geonoma polyandra , overlooked the earlier name of G. multisecta . This was understandable given that the type specimen of G. multisecta lacked staminate flowers. However, this specimen, with its veins raised and rectangular in cross-section adaxially and leaf and inflorescence morphology certainly represents the earliest name for this species. Geonoma multisecta is a member of a group of related species including G. macrostachys , G. paradoxa , G. poiteauana , and G. schizocarpa , and is the only one of which to have more than six stamens. Subspecific variation:— No trait varies within this species, nor is there any geographic disjunction.