Revision of the family Felixaraeidae (Scleractinia; Cretaceous)
Author
Löser, Hannes
text
Geodiversitas
2013
2013-12-27
35
4
747
765
http://dx.doi.org/10.5252/g2013n4a1
journal article
10.5252/g2013n4a1
1638-9395
5376738
Family
FELIXARAEIDAE
Beauvais, 1982
DIAGNOSIS. — Corals of various organisation
types
(astreoid, phaceloid, solitary, thamnasterioid) with generally large calices (>
5 mm
). Radial elements composed of large trabeculae, with very thick septa in the first septal generations. Radial elements perforated, mainly in septa of younger generations. Septal upper border with coarse granules, septal lateral faces smooth or with thorns. Costae present. Synapticulae abundant and strong. Endotheca varies from genus to genus; it may be absent or very well developed. Columella generally present, parietal.
GENERA OF THE FAMILY. —
Brachycaulia
Beauvais, 1982
,
Cretastraea
Kühn
in Kühn & Andrusov, 1930 (junior objective synonym of
Pseudofavia
),
Felixaraea
Beauvais, 1982
,
Filkornia
Löser, 2012
,
Marcelohelia
n. gen.
,
Paracycloseris
Wells, 1934
,
Pseudofavia
Oppenheim, 1930
,
Thecoseriopsis
Alloiteau, 1952
,
Trechmannaria
Wells, 1935
.
RANGE. — Late Turonian to?Danian. The first indication (
Pseudofavia paronai
), dated as Turonian, is not well constrained. The last occurrence is similarly not well constrained; the family ranges into the Maastrichtian, but extension into the Danian is questionable.
SYSTEMATIC POSITION
The position of the family within the
Fungiina
is questionable. As mentioned by
Morycowa & Roniewicz (1995)
, the suborder
Fungiina
collects different families that are characterized only by porous radial elements and the presence of synapticulae.
Fungiina
sensu stricto are corals with synapticular bars (fulturae aucct.), which are found, for instance, in the
Asteroseriidae
and
Fungiidae
. Synapticular bars are absent in
Felixaraeidae
, and therefore the position of this family within the suborder
Fungiina
is preliminary.
Baron-Szabo (2008) included the family in the synonymy of the family
Haplaraeidae
. The
Haplaraeidae
differs from the
Felixaraeidae
by having radial elements that are very regular in thickness (all septal generations have the same thickness) and only perforated at their inner margins (based on observations of the
type
of the
type
species,
Haplaraea elegans
Milaschewitsch, 1876
, SMNS 21874).
COMPARISON
Very strong radial elements and the presence of synapticulae are also observed in the Late Cretaceous family
Lamellofungiidae Alloiteau, 1952
. The family currently encompasses only the
type
genus
Lamellofungia
Alloiteau, 1952
. The genus
Lamnastrea
Reig-Oriol, 1997
may also belong to this family. The members of this family differ from those of the
Felixaraeidae
by having radial elements with spare pores at the inner margins.
REMARKS
The porosity of the radial elements appears irregular. It seem that both, older and younger septal generations are regularly perforated but the size of pores may vary within the same specimen resulting that almost compact to regularly perforated radial elements occur together. This feature can be observed in almost all specimens. Only
Filkornia
seems to have more compact septa.