Species delimitation of colour polymorphic Cladophorus (Coleoptera: Lycidae) from New Guinea Author Bocak, Ladislav text Zootaxa 2017 2017-09-18 4320 3 505 522 journal article 32076 10.11646/zootaxa.4320.3.6 e80fc464-0c2a-4003-a867-ab8b5b3b399e 1175-5326 1051951 Ca50E5A5-B417-4000-B67D-5F6028Da9154 Cladophorus humeralis Kleine, 1926 ( Figs. 10, 24 , 40 , 61–62 ) Cladophorus humeralis Kleine, 1926 : 130 . Type material. ‘ Co-Typus’ [ syntype , No. 274]. Male , New Guinea , Bomberai Peninsula , Kapaur ( MIZW ). Other material examined. Male, female . Indonesia , West Papua prov., DMP-Wasior, 100–200 m (A00434–35, LMBC ). Diagnosis. C. humeralis has a unique elytral colour pattern with brown humeri and a longitudinal patch in the posterior part of each elytron ( Fig. 40 ). Further, the phallus of this species is robust and gradually acuminate apically ( Figs. 61–62 ). C. manokwarensis was identified as a sister species of C. humeralis in the molecular phylogeny ( Fig. 4 ). Although these species are very close each to other, they differ besides the colour pattern in the shape of the antennomere 3 and the relative size of eyes. The shape of phallus differs only slightly in this pair of species ( Figs. 61–62, 65–66 ) Redescription. Male. Body large, 15.5–15.7 mm long, dark brown to black including pronotum and substantial part of elytra ( Fig. 40 ), humeri light brown, bright humeral patches separated by dark coloured area along inner margin of elytra; additionally long, light patch in posterior part of each elytron, posterior patch with gradual transition between dark and light coloured areas of elytron ( Fig. 40 ). Head small, with hemispherically prominent eyes, frontal interocular distance 1.92 times maximum diameter; antennae long, reaching beyond two thirds of elytral length, flabellate, antennomere 3 with lamella 2.55 times longer than stem, lamella attached to apical part of antennomere ( Fig. 10 ). Pronotum transverse, 1.43 times wider than long at midline, lateral margins apparently elevated, frontal margin widely rounded, frontal angles inconspicuous, rounded, lateral margins slightly convex, posterior angles prominent, all carinae well-developed ( Fig. 24 ). Elytra flat, slender, subparallel-sided, transverse costae well-developed, regular, elytral areoles slightly transverse ( Fig. 40 ). Phallus robust, parallel-sided in middle, with gradually acuminate apex ( Figs. 61–62 ). Measurements. BL 15.5–15.7 mm , PL 2.10 mm , PW 3.00– 3.05 mm , WH 1.70–1.75 mm , Edist 1.15 mm , Ediam 0.60 mm , LE 13.2 mm , antennomere 3: stem 1.0 mm, lamella 2.55 mm . Distribution. Western New Guinea , Bomberai Peninsula, Teluk Wondama regency ( Fig. 3 )